中国水稻科学 ›› 2020, Vol. 34 ›› Issue (1): 17-27.DOI: 10.16819/j.1001-7216.2020.9052
王小雷, 刘杨, 孙晓棠, 欧阳林娟, 潘锦龙, 彭小松, 陈小荣, 贺晓鹏, 傅军如, 边建民, 胡丽芳, 徐杰, 贺浩华*(), 朱昌兰*()
收稿日期:
2019-04-30
修回日期:
2019-06-13
出版日期:
2020-01-10
发布日期:
2020-01-10
通讯作者:
贺浩华,朱昌兰
作者简介:
作者简介:#共同第一作者
基金资助:
Xiaolei WANG, Yang LIU, Xiaotang SUN, Linjuan OUYANG, Jinlong PAN, Xiaosong PENG, Xiaorong CHEN, Xiaopeng HE, Junru FU, Jianmin BIAN, Lifang HU, Jie XU, Haohua HE*(), Changlan ZHU*()
Received:
2019-04-30
Revised:
2019-06-13
Online:
2020-01-10
Published:
2020-01-10
Contact:
Haohua HE, Changlan ZHU
About author:
About author:#These authors contributed equally to this work
摘要:
【目的】挖掘新的稻米品质性状QTL并利用分子育种技术改良稻米品质。【方法】利用构建的一套以籼稻香型恢复系昌恢121为背景亲本,以优质粳稻越光为供体亲本的染色体片段代换系(CSSLs)为材料,在4个环境下对稻米品质性状进行QTL检测及稳定性分析。【结果】在4个环境下共检测到44个QTL,其中6个QTL能在多个环境下被检测到;第2、3、5、6和10染色体上存在多效QTL簇,对稻米品质性状具有明显的调控作用;第1、6和12染色体上7个QTL能在不同环境下稳定表达。【结论】第1染色体RM3143-RM1117区间qPGWC1和第12染色体RM3331-RM5479区间qPaT12是两个新的稳定表达QTL。
中图分类号:
王小雷, 刘杨, 孙晓棠, 欧阳林娟, 潘锦龙, 彭小松, 陈小荣, 贺晓鹏, 傅军如, 边建民, 胡丽芳, 徐杰, 贺浩华, 朱昌兰. 不同环境下稻米品质性状QTL的检测及稳定性分析[J]. 中国水稻科学, 2020, 34(1): 17-27.
Xiaolei WANG, Yang LIU, Xiaotang SUN, Linjuan OUYANG, Jinlong PAN, Xiaosong PENG, Xiaorong CHEN, Xiaopeng HE, Junru FU, Jianmin BIAN, Lifang HU, Jie XU, Haohua HE, Changlan ZHU. Identification and Stability Analysis of QTL for Grain Quality Traits Under Multiple Environments in Rice[J]. Chinese Journal OF Rice Science, 2020, 34(1): 17-27.
性状 Trait | 环境 E | 亲本 Parent | CSSL群体 CSSL population | ||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|
昌恢121 Changhui 121 | 越光 Koshihikari | 变异范围 Range | 平均±标准差 Mean±SD | 峰度 Kurtosis | 偏度 Skewness | ||||||||
垩白粒率PGWC/% | E1 | 4.5±0.5 | 5.0±0.0 | 4.0~42.0 | 14.9±9.6 | 1.4 | 1.4 | ||||||
E2 | 2.5±0.5 | 3.0±1.0 | 0.5~49.0 | 6.2±8.1 | 13.8 | 3.5 | |||||||
E3 | 3.5±1.5 | 33.0±2.0 | 2.0~56.0 | 16.0±9.8 | 3.4 | 1.3 | |||||||
E4 | 7.0±0.5 | 6.0±1.0 | 4.0~41.0 | 15.8±7.8 | 1.2 | 1.1 | |||||||
胶稠度 GC/mm | E1 | 30.0±2.0 | 51.5±0.5 | 24.5~66.0 | 36.0±9.8 | 2.4 | 1.7 | ||||||
E2 | 30.5±0.5 | 56.5±0.5 | 30.5~92.5 | 43.5±15.9 | 2.4 | 1.7 | |||||||
E3 | 55.5±1.5 | 82.5±1.5 | 34.0~87.5 | 57.4±13.3 | -0.7 | 0.3 | |||||||
E4 | 32.5±1.5 | 67.5±1.5 | 31.5~100.0 | 59.3±19.0 | -0.6 | 0.4 | |||||||
蛋白质含量PC/% | E1 | 8.6±0.2 | 8.11±0.1 | 5.9~10.6 | 7.7±0.8 | 1.6 | 0.8 | ||||||
E2 | 8.9±0.1 | 8.98±0.1 | 6.4~10.1 | 8.5±0.8 | 0.8 | 0.7 | |||||||
E3 | 10.2±0.0 | 9.04±0.0 | 70~11.3 | 8.6±1.0 | 0.1 | 0.6 | |||||||
E4 | 7.1±0.0 | 7.44±0.0 | 5.6~7.9 | 6.5±0.5 | 0.2 | 0.7 | |||||||
最高黏度PKV/cP | E1 | 4936.0±91.0 | 5718.0±29.0 | 3850.5~5922.0 | 4962.1±434.3 | 0.6 | -0.3 | ||||||
E2 | 4972.0±24.0 | 4557.5±77.0 | 3193.5~6027.0 | 4387.6±536.1 | 0.7 | 0.7 | |||||||
E3 | 3900.0±22.0 | 4448.0±73.0 | 2567.0~4949.5 | 4152.1±336.9 | 8.0 | -1.6 | |||||||
E4 | 4878.0±82.0 | 4944.5±48.5 | 4015.0~5194.5 | 4462.5±228.8 | 0.7 | 0.6 | |||||||
热浆黏度HPV/cP | E1 | 3412.5±100.5 | 2321.5±56.5 | 2163.5~4098.5 | 3298.6±456.0 | 0.3 | -1.1 | ||||||
E2 | 4136.5±54.5 | 2965.5±50.5 | 2637.5~4645.0 | 3485.9±431.3 | 0.1 | 0.2 | |||||||
E3 | 2784.0±86.0 | 1948.0±52.0 | 1592.5~3709.5 | 2807.3±428.7 | 1.6 | -1.2 | |||||||
E4 | 3404.5±48.5 | 2818.5±55.5 | 2029.0~3589.0 | 3097.8±340.4 | 2.2 | -1.5 | |||||||
崩解值BDV/cP | E1 | 1523.5±103.5 | 3396.5±98.5 | 1037.0~3713.5 | 1663.5±620.4 | 3.5 | 2.1 | ||||||
E2 | 835.5±30.5 | 1592.0±27.0 | 298.0~3000.0 | 901.7±490.8 | 5.5 | 2.2 | |||||||
E3 | 1116.0±60.7 | 2500.0±17.0 | 700.5~2556.0 | 1344.8±354.2 | 2.7 | 1.4 | |||||||
E4 | 1473.5±75.5 | 2126.0±81.0 | 903.0~3165.5 | 1364.8±436.2 | 6.2 | 2.4 | |||||||
冷浆黏度CPV/cP | E1 | 6664.0±34.5 | 3723.5±18.5 | 3351.0~7426.0 | 6008.0±1028.7 | 1.4 | -1.5 | ||||||
E2 | 7332.5±4.5 | 4775.5±40.5 | 3963.0~8000.0 | 6459.8±951.6 | 0.4 | -1.0 | |||||||
E3 | 5643.5±47.5 | 2901.5±6.5 | 2790.5~7454.5 | 5350.3±870.2 | 2.7 | -1.5 | |||||||
E4 | 6222.0±35.0 | 4224.5±37.5 | 3254.0~6477.5 | 5641.7±773.1 | 3.1 | -2.0 | |||||||
消减值SBV/cP | E1 | 1728.0±36.0 | -1994.5±97.5 | -2505.0~2160.5 | 1045.9±1177.2 | 3.1 | -2.1 | ||||||
E2 | 2360.5±28.5 | 218.0±37.0 | -1765.0~3005.5 | 2072.3±982.7 | 4.6 | -2.3 | |||||||
E3 | 1743.5±31.5 | -1546.5±12.5 | -1434.5~2505.0 | 1198.3±760.4 | 3.9 | -2.1 | |||||||
E4 | 1344.0±42.0 | -720.0±31.0 | -1940.5~2189.5 | 1179.2±865.0 | 4.6 | -2.3 | |||||||
回复值CSV/cP | E1 | 3251.5±125.5 | 1402.0±20.0 | 1187.5~3546.5 | 2709.4±602.8 | 1.9 | -1.7 | ||||||
E2 | 3196.0±59.0 | 1810.0±10.0 | 1235.0~3757.5 | 2973.9±584.8 | 1.5 | -1.5 | |||||||
E3 | 2859.5±27.5 | 953.5±5.5 | 1121.5~3745.0 | 2543.1±545.3 | 1.0 | -1.1 | |||||||
E4 | 2817.5±55.5 | 1406.0±28.0 | 1138.5~3460.0 | 2544.0±520.0 | 1.2 | -1.2 | |||||||
成糊温度PaT/℃ | E1 | 72.4±0.3 | 74.6±0.7 | 68.1~76.0 | 71.8±1.5 | 1.3 | 0.3 | ||||||
E2 | 78.4±0.8 | 80.7±0.1 | 74.1~87.0 | 81.1±3.3 | -0.6 | -0.6 | |||||||
E3 | 70.6±0.1 | 76.8±0.3 | 70.0~76.4 | 72.1±1.2 | 1.2 | 0.9 | |||||||
E4 | 72.1±0.1 | 73.7±0.3 | 70.9~74.8 | 72.1±1.5 | 1.6 | 1.1 | |||||||
成糊时间PeT/min | E1 | 5.9±0.1 | 5.7±0.1 | 5.5~6.2 | 5.9±0.1 | 1.4 | -1.0 | ||||||
E2 | 6.1±0.0 | 6.0±0.0 | 5.7~6.4 | 6.0±0.1 | 0.4 | 0.2 | |||||||
E3 | 6.0±0.1 | 5.7±0.3 | 5.7~6.7 | 6.0±0.2 | 0.3 | 0.6 | |||||||
E4 | 6.0±0.2 | 6.2±0.1 | 5.6~6.4 | 6.1±0.2 | -0.2 | -0.2 |
表1 亲本及CSSLs群体在4个环境中的稻米品质性状
Table 1 Phenotypic performance of CSSLs population and its parents in 4 environments.
性状 Trait | 环境 E | 亲本 Parent | CSSL群体 CSSL population | ||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|
昌恢121 Changhui 121 | 越光 Koshihikari | 变异范围 Range | 平均±标准差 Mean±SD | 峰度 Kurtosis | 偏度 Skewness | ||||||||
垩白粒率PGWC/% | E1 | 4.5±0.5 | 5.0±0.0 | 4.0~42.0 | 14.9±9.6 | 1.4 | 1.4 | ||||||
E2 | 2.5±0.5 | 3.0±1.0 | 0.5~49.0 | 6.2±8.1 | 13.8 | 3.5 | |||||||
E3 | 3.5±1.5 | 33.0±2.0 | 2.0~56.0 | 16.0±9.8 | 3.4 | 1.3 | |||||||
E4 | 7.0±0.5 | 6.0±1.0 | 4.0~41.0 | 15.8±7.8 | 1.2 | 1.1 | |||||||
胶稠度 GC/mm | E1 | 30.0±2.0 | 51.5±0.5 | 24.5~66.0 | 36.0±9.8 | 2.4 | 1.7 | ||||||
E2 | 30.5±0.5 | 56.5±0.5 | 30.5~92.5 | 43.5±15.9 | 2.4 | 1.7 | |||||||
E3 | 55.5±1.5 | 82.5±1.5 | 34.0~87.5 | 57.4±13.3 | -0.7 | 0.3 | |||||||
E4 | 32.5±1.5 | 67.5±1.5 | 31.5~100.0 | 59.3±19.0 | -0.6 | 0.4 | |||||||
蛋白质含量PC/% | E1 | 8.6±0.2 | 8.11±0.1 | 5.9~10.6 | 7.7±0.8 | 1.6 | 0.8 | ||||||
E2 | 8.9±0.1 | 8.98±0.1 | 6.4~10.1 | 8.5±0.8 | 0.8 | 0.7 | |||||||
E3 | 10.2±0.0 | 9.04±0.0 | 70~11.3 | 8.6±1.0 | 0.1 | 0.6 | |||||||
E4 | 7.1±0.0 | 7.44±0.0 | 5.6~7.9 | 6.5±0.5 | 0.2 | 0.7 | |||||||
最高黏度PKV/cP | E1 | 4936.0±91.0 | 5718.0±29.0 | 3850.5~5922.0 | 4962.1±434.3 | 0.6 | -0.3 | ||||||
E2 | 4972.0±24.0 | 4557.5±77.0 | 3193.5~6027.0 | 4387.6±536.1 | 0.7 | 0.7 | |||||||
E3 | 3900.0±22.0 | 4448.0±73.0 | 2567.0~4949.5 | 4152.1±336.9 | 8.0 | -1.6 | |||||||
E4 | 4878.0±82.0 | 4944.5±48.5 | 4015.0~5194.5 | 4462.5±228.8 | 0.7 | 0.6 | |||||||
热浆黏度HPV/cP | E1 | 3412.5±100.5 | 2321.5±56.5 | 2163.5~4098.5 | 3298.6±456.0 | 0.3 | -1.1 | ||||||
E2 | 4136.5±54.5 | 2965.5±50.5 | 2637.5~4645.0 | 3485.9±431.3 | 0.1 | 0.2 | |||||||
E3 | 2784.0±86.0 | 1948.0±52.0 | 1592.5~3709.5 | 2807.3±428.7 | 1.6 | -1.2 | |||||||
E4 | 3404.5±48.5 | 2818.5±55.5 | 2029.0~3589.0 | 3097.8±340.4 | 2.2 | -1.5 | |||||||
崩解值BDV/cP | E1 | 1523.5±103.5 | 3396.5±98.5 | 1037.0~3713.5 | 1663.5±620.4 | 3.5 | 2.1 | ||||||
E2 | 835.5±30.5 | 1592.0±27.0 | 298.0~3000.0 | 901.7±490.8 | 5.5 | 2.2 | |||||||
E3 | 1116.0±60.7 | 2500.0±17.0 | 700.5~2556.0 | 1344.8±354.2 | 2.7 | 1.4 | |||||||
E4 | 1473.5±75.5 | 2126.0±81.0 | 903.0~3165.5 | 1364.8±436.2 | 6.2 | 2.4 | |||||||
冷浆黏度CPV/cP | E1 | 6664.0±34.5 | 3723.5±18.5 | 3351.0~7426.0 | 6008.0±1028.7 | 1.4 | -1.5 | ||||||
E2 | 7332.5±4.5 | 4775.5±40.5 | 3963.0~8000.0 | 6459.8±951.6 | 0.4 | -1.0 | |||||||
E3 | 5643.5±47.5 | 2901.5±6.5 | 2790.5~7454.5 | 5350.3±870.2 | 2.7 | -1.5 | |||||||
E4 | 6222.0±35.0 | 4224.5±37.5 | 3254.0~6477.5 | 5641.7±773.1 | 3.1 | -2.0 | |||||||
消减值SBV/cP | E1 | 1728.0±36.0 | -1994.5±97.5 | -2505.0~2160.5 | 1045.9±1177.2 | 3.1 | -2.1 | ||||||
E2 | 2360.5±28.5 | 218.0±37.0 | -1765.0~3005.5 | 2072.3±982.7 | 4.6 | -2.3 | |||||||
E3 | 1743.5±31.5 | -1546.5±12.5 | -1434.5~2505.0 | 1198.3±760.4 | 3.9 | -2.1 | |||||||
E4 | 1344.0±42.0 | -720.0±31.0 | -1940.5~2189.5 | 1179.2±865.0 | 4.6 | -2.3 | |||||||
回复值CSV/cP | E1 | 3251.5±125.5 | 1402.0±20.0 | 1187.5~3546.5 | 2709.4±602.8 | 1.9 | -1.7 | ||||||
E2 | 3196.0±59.0 | 1810.0±10.0 | 1235.0~3757.5 | 2973.9±584.8 | 1.5 | -1.5 | |||||||
E3 | 2859.5±27.5 | 953.5±5.5 | 1121.5~3745.0 | 2543.1±545.3 | 1.0 | -1.1 | |||||||
E4 | 2817.5±55.5 | 1406.0±28.0 | 1138.5~3460.0 | 2544.0±520.0 | 1.2 | -1.2 | |||||||
成糊温度PaT/℃ | E1 | 72.4±0.3 | 74.6±0.7 | 68.1~76.0 | 71.8±1.5 | 1.3 | 0.3 | ||||||
E2 | 78.4±0.8 | 80.7±0.1 | 74.1~87.0 | 81.1±3.3 | -0.6 | -0.6 | |||||||
E3 | 70.6±0.1 | 76.8±0.3 | 70.0~76.4 | 72.1±1.2 | 1.2 | 0.9 | |||||||
E4 | 72.1±0.1 | 73.7±0.3 | 70.9~74.8 | 72.1±1.5 | 1.6 | 1.1 | |||||||
成糊时间PeT/min | E1 | 5.9±0.1 | 5.7±0.1 | 5.5~6.2 | 5.9±0.1 | 1.4 | -1.0 | ||||||
E2 | 6.1±0.0 | 6.0±0.0 | 5.7~6.4 | 6.0±0.1 | 0.4 | 0.2 | |||||||
E3 | 6.0±0.1 | 5.7±0.3 | 5.7~6.7 | 6.0±0.2 | 0.3 | 0.6 | |||||||
E4 | 6.0±0.2 | 6.2±0.1 | 5.6~6.4 | 6.1±0.2 | -0.2 | -0.2 |
性状Trait | 环境E | 性状Trait | ||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|
PGWC | GC | PC | PKV | HPV | BDV | CPV | SBV | CSV | PaT | |||
GC | E1 | 0.08 | ||||||||||
E2 | 0.11 | |||||||||||
E3 | 0.05 | |||||||||||
E4 | –0.14 | |||||||||||
PC | E1 | –0.08 | –0.13 | |||||||||
E2 | –0.31* | –0.00 | ||||||||||
E3 | –0.42** | –0.29* | ||||||||||
E4 | –0.18 | –0.05 | ||||||||||
PKV | E1 | –0.18 | 0.27* | –0.21 | ||||||||
E2 | 0.23 | 0.25 | –0.76** | |||||||||
E3 | 0.02 | 0.19 | –0.31* | |||||||||
E4 | –0.07 | 0.37** | –0.03 | |||||||||
HPV | E1 | –0.04 | –0.48** | –0.08 | 0.03 | |||||||
E2 | –0.23 | –0.48** | –0.46** | 0.50** | ||||||||
E3 | 0.25 | –0.27* | –0.28* | 0.55** | ||||||||
E4 | 0.16 | –0.35** | –0.08 | –0.14 | ||||||||
BDV | E1 | –0.09 | 0.54** | –0.09 | 0.68** | –0.71** | ||||||
E2 | 0.45** | 0.70** | –0.43** | 0.65** | –0.33* | |||||||
E3 | –0.27* | 0.48** | 0.04 | 0.26 | –0.66** | |||||||
E4 | –0.16 | 0.47** | 0.05 | 0.64** | –0.86** | |||||||
CPV | E1 | –0.04 | –0.50** | –0.05 | –0.16 | 0.96** | –0.82** | |||||
E2 | –0.34** | –0.62** | –0.33** | 0.22 | 0.91** | –0.56** | ||||||
E3 | 0.17 | –0.41** | –0.15 | 0.44** | 0.85** | –0.59** | ||||||
E4 | –0.03 | –0.47** | –0.05 | –0.28* | 0.84** | –0.80** | ||||||
SBV | E1 | 0.03 | –0.54** | 0.03 | –0.51** | 0.83** | –0.96** | 0.93** | ||||
E2 | –0.45** | –0.74** | 0.10 | –0.33** | 0.61** | –0.90** | 0.85** | |||||
E3 | 0.19 | –0.54** | –0.04 | 0.06 | 0.71** | –0.77** | 0.92** | |||||
E4 | –0.01 | –0.51** | –0.04 | –0.51** | 0.79** | –0.89** | 0.97** | |||||
CSV | E1 | –0.04 | –0.49** | –0.03 | –0.29* | 0.89** | –0.85** | 0.98** | 0.96** | |||
E2 | –0.38** | –0.65** | –0.19 | –0.01 | 0.75** | –0.67** | 0.95** | 0.93** | ||||
E3 | 0.08 | –0.43** | –0.03 | 0.26 | 0.55** | –0.41** | 0.91** | 0.90** | ||||
E4 | –0.15 | –0.46** | –0.03 | –0.32* | 0.60** | –0.63** | 0.94** | 0.92** | ||||
PaT | E1 | –0.18 | 0.16 | 0.48** | 0.16 | –0.54** | 0.51** | –0.57** | –0.55** | –0.56** | ||
E2 | –0.29* | –0.11 | 0.76** | –0.89** | –0.45** | –0.58** | –0.26* | 0.23 | –0.09 | |||
E3 | –0.05 | –0.18 | 0.27* | –0.42** | –0.40** | 0.09 | –0.43** | –0.30* | –0.36** | |||
E4 | –0.19 | –0.01 | 0.44** | –0.14 | –0.32** | 0.18 | –0.23 | –0.17 | –0.14 | |||
PeT | E1 | 0.18 | –0.37** | 0.14 | –0.38** | 0.67** | –0.76** | 0.66** | 0.71** | 0.61** | –0.30* | |
E2 | –0.18 | –0.51** | 0.48** | –0.44** | 0.21 | –0.67** | 0.24 | 0.47** | 0.24 | 0.49** | ||
E3 | 0.21 | –0.10 | –0.13 | –0.06 | 0.55** | –0.69** | 0.17 | 0.21 | –0.17 | 0.05 | ||
E4 | 0.37** | –0.18 | –0.04 | –0.32* | 0.64** | –0.67** | 0.23 | 0.29* | –0.07 | –0.05 |
表2 昌恢121/越光CSSLs群体在4个环境中稻米各品质性状间的相关系数
Table 2 Coefficients of pairwise correlations among rice quality traits in the Changhui 121/Koshihikari CSSLs population.
性状Trait | 环境E | 性状Trait | ||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|
PGWC | GC | PC | PKV | HPV | BDV | CPV | SBV | CSV | PaT | |||
GC | E1 | 0.08 | ||||||||||
E2 | 0.11 | |||||||||||
E3 | 0.05 | |||||||||||
E4 | –0.14 | |||||||||||
PC | E1 | –0.08 | –0.13 | |||||||||
E2 | –0.31* | –0.00 | ||||||||||
E3 | –0.42** | –0.29* | ||||||||||
E4 | –0.18 | –0.05 | ||||||||||
PKV | E1 | –0.18 | 0.27* | –0.21 | ||||||||
E2 | 0.23 | 0.25 | –0.76** | |||||||||
E3 | 0.02 | 0.19 | –0.31* | |||||||||
E4 | –0.07 | 0.37** | –0.03 | |||||||||
HPV | E1 | –0.04 | –0.48** | –0.08 | 0.03 | |||||||
E2 | –0.23 | –0.48** | –0.46** | 0.50** | ||||||||
E3 | 0.25 | –0.27* | –0.28* | 0.55** | ||||||||
E4 | 0.16 | –0.35** | –0.08 | –0.14 | ||||||||
BDV | E1 | –0.09 | 0.54** | –0.09 | 0.68** | –0.71** | ||||||
E2 | 0.45** | 0.70** | –0.43** | 0.65** | –0.33* | |||||||
E3 | –0.27* | 0.48** | 0.04 | 0.26 | –0.66** | |||||||
E4 | –0.16 | 0.47** | 0.05 | 0.64** | –0.86** | |||||||
CPV | E1 | –0.04 | –0.50** | –0.05 | –0.16 | 0.96** | –0.82** | |||||
E2 | –0.34** | –0.62** | –0.33** | 0.22 | 0.91** | –0.56** | ||||||
E3 | 0.17 | –0.41** | –0.15 | 0.44** | 0.85** | –0.59** | ||||||
E4 | –0.03 | –0.47** | –0.05 | –0.28* | 0.84** | –0.80** | ||||||
SBV | E1 | 0.03 | –0.54** | 0.03 | –0.51** | 0.83** | –0.96** | 0.93** | ||||
E2 | –0.45** | –0.74** | 0.10 | –0.33** | 0.61** | –0.90** | 0.85** | |||||
E3 | 0.19 | –0.54** | –0.04 | 0.06 | 0.71** | –0.77** | 0.92** | |||||
E4 | –0.01 | –0.51** | –0.04 | –0.51** | 0.79** | –0.89** | 0.97** | |||||
CSV | E1 | –0.04 | –0.49** | –0.03 | –0.29* | 0.89** | –0.85** | 0.98** | 0.96** | |||
E2 | –0.38** | –0.65** | –0.19 | –0.01 | 0.75** | –0.67** | 0.95** | 0.93** | ||||
E3 | 0.08 | –0.43** | –0.03 | 0.26 | 0.55** | –0.41** | 0.91** | 0.90** | ||||
E4 | –0.15 | –0.46** | –0.03 | –0.32* | 0.60** | –0.63** | 0.94** | 0.92** | ||||
PaT | E1 | –0.18 | 0.16 | 0.48** | 0.16 | –0.54** | 0.51** | –0.57** | –0.55** | –0.56** | ||
E2 | –0.29* | –0.11 | 0.76** | –0.89** | –0.45** | –0.58** | –0.26* | 0.23 | –0.09 | |||
E3 | –0.05 | –0.18 | 0.27* | –0.42** | –0.40** | 0.09 | –0.43** | –0.30* | –0.36** | |||
E4 | –0.19 | –0.01 | 0.44** | –0.14 | –0.32** | 0.18 | –0.23 | –0.17 | –0.14 | |||
PeT | E1 | 0.18 | –0.37** | 0.14 | –0.38** | 0.67** | –0.76** | 0.66** | 0.71** | 0.61** | –0.30* | |
E2 | –0.18 | –0.51** | 0.48** | –0.44** | 0.21 | –0.67** | 0.24 | 0.47** | 0.24 | 0.49** | ||
E3 | 0.21 | –0.10 | –0.13 | –0.06 | 0.55** | –0.69** | 0.17 | 0.21 | –0.17 | 0.05 | ||
E4 | 0.37** | –0.18 | –0.04 | –0.32* | 0.64** | –0.67** | 0.23 | 0.29* | –0.07 | –0.05 |
位点 QTL | 区间 Marker interval | LOD值LOD value | 贡献率PVE/% | 加性效应Additive effect | |||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|
E1 | E2 | E3 | E4 | E1 | E2 | E3 | E4 | E1 | E2 | E3 | E4 | ||
qPGWC1 | RM3143-RM1117 | 5.02 | 10.45 | 4.40 | 29.34 | 57.67 | 30.84 | 18.40 | 13.34 | 20.36 | |||
qPGWC2 | RM5529-RM7581 | 4.27 | 16.61 | 4.58 | |||||||||
qPGWC5 | RM3345-RM17954 | 2.93 | 10.99 | 8.89 | |||||||||
qPGWC6 | RM7088-RM7311 | 3.54 | 19.27 | 10.65 | |||||||||
qGC5 | RM3345-RM17954 | 6.54 | 34.67 | 15.72 | |||||||||
qGC6.1 | RM1369-RM510 | 6.76 | 5.64 | 41.02 | 25.07 | 15.78 | 13.60 | ||||||
qGC6.2 | RM510-RM1163 | 2.99 | 22.48 | 10.86 | |||||||||
qGC9 | STS-P0692F07- STS-OJ1001_G09 | 3.48 | 16.05 | 6.97 | |||||||||
qPC10.1 | RM6737-RM25923 | 2.71 | 19.04 | ﹣0.59 | |||||||||
qPC10.2 | RM25923-RM271 | 2.76 | 13.15 | 0.28 |
表3 昌恢121/越光CSSLs群体垩白粒率、胶稠度和蛋白质含量QTL分析
Table 3 QTL detected for PGWC, GC, PC in the CSSLs population.
位点 QTL | 区间 Marker interval | LOD值LOD value | 贡献率PVE/% | 加性效应Additive effect | |||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|
E1 | E2 | E3 | E4 | E1 | E2 | E3 | E4 | E1 | E2 | E3 | E4 | ||
qPGWC1 | RM3143-RM1117 | 5.02 | 10.45 | 4.40 | 29.34 | 57.67 | 30.84 | 18.40 | 13.34 | 20.36 | |||
qPGWC2 | RM5529-RM7581 | 4.27 | 16.61 | 4.58 | |||||||||
qPGWC5 | RM3345-RM17954 | 2.93 | 10.99 | 8.89 | |||||||||
qPGWC6 | RM7088-RM7311 | 3.54 | 19.27 | 10.65 | |||||||||
qGC5 | RM3345-RM17954 | 6.54 | 34.67 | 15.72 | |||||||||
qGC6.1 | RM1369-RM510 | 6.76 | 5.64 | 41.02 | 25.07 | 15.78 | 13.60 | ||||||
qGC6.2 | RM510-RM1163 | 2.99 | 22.48 | 10.86 | |||||||||
qGC9 | STS-P0692F07- STS-OJ1001_G09 | 3.48 | 16.05 | 6.97 | |||||||||
qPC10.1 | RM6737-RM25923 | 2.71 | 19.04 | ﹣0.59 | |||||||||
qPC10.2 | RM25923-RM271 | 2.76 | 13.15 | 0.28 |
位点 QTL | 区间 Marker interval | LOD值LOD value | 贡献率PVE/% | ||||||
---|---|---|---|---|---|---|---|---|---|
E1 | E2 | E3 | E4 | E1 | E2 | E3 | E4 | ||
qPKV1 | RM6292-RM5362 | 3.08 | 10.44 | ||||||
qPKV3 | RM6297-RM1278 | 6.30 | 40.98 | ||||||
qPKV6.1 | RM1369-RM510 | 3.42 | 25.32 | ||||||
qPKV6.2 | RM510-RM1163 | 5.61 | 20.35 | ||||||
qPKV10 | RM6737-RM25923 | 4.19 | 22.55 | ||||||
qPKV11 | STS-OSJNBa0095P22-RM5766 | 3.05 | 15.69 | ||||||
qHPV3 | RM6297-RM1278 | 5.61 | 17.01 | ||||||
qHPV6 | RM1369-RM510 | 12.22 | 4.98 | 12.64 | 30.11 | 64.75 | 30.21 | 53.41 | 76.71 |
qHPV8 | RM6838-RM5767 | 2.79 | 3.21 | ||||||
qBDV3 | RM15887-RM15903 | 3.79 | 3.34 | ||||||
qBDV5 | RM3345-RM17954 | 3.72 | 14.25 | ||||||
qBDV6 | RM1369-RM510 | 25.94 | 4.98 | 11.39 | 35.00 | 89.05 | 32.21 | 61.47 | 77.94 |
qCPV3.1 | RM15887-RM15903 | 2.78 | 4.40 | ||||||
qCPV3.2 | RM6297-RM1278 | 11.11 | 13.98 | ||||||
qCPV6 | RM1369-RM510 | 20.80 | 10.46 | 22.26 | 45.45 | 80.40 | 50.81 | 60.34 | 90.23 |
qCPV8 | RM5556-RM6838 | 11.58 | 14.92 | ||||||
qSBV2 | RM5529-RM7581 | 2.88 | 5.12 | ||||||
qSBV3 | RM15887-RM15903 | 5.06 | 1.61 | ||||||
qSBV5 | RM3345-RM17954 | 3.54 | 6.45 | ||||||
qSBV6.1 | RM1369-RM510 | 33.68 | 19.23 | 22.89 | 56.05 | 94.34 | 70.67 | 85.28 | 90.49 |
qSBV6.2 | RM510-RM1163 | 5.81 | 1.89 | ||||||
qSBV8 | RM7049-RM7556 | 3.37 | 1.03 | ||||||
qCSV3 | RM15887-RM15903 | 3.23 | 3.72 | ||||||
qCSV5 | RM3476-RM3790 | 2.74 | 2.97 | ||||||
qCSV6 | RM1369-RM510 | 24.77 | 12.92 | 11.12 | 30.99 | 85.33 | 63.53 | 60.61 | 76.79 |
qPaT4 | RM3839-RM303 | 3.53 | 5.04 | ||||||
qPaT6.1 | RM1369-RM510 | 2.70 | 11.27 | 15.05 | 19.62 | ||||
qPaT6.2 | RM510-RM1163 | 4.63 | 32.63 | ||||||
qPaT10 | RM6737-RM25923 | 4.59 | 30.13 | ||||||
qPaT12 | RM3331-RM5479 | 3.90 | 20.99 | 22.88 | 48.48 | ||||
qPeT6 | RM1369-RM510 | 11.05 | 3.59 | 48.48 | 16.77 | ||||
qPeT7.1 | RM1132-RM8261 | 3.51 | 10.78 | ||||||
qPeT7.2 | STS-OJ1123_C12-RM5752 | 4.47 | 23.25 | ||||||
qPeT7.3 | RM7273-RM3404 | 2.95 | 14.44 |
表4 昌恢121/越光CSSLs群体8个RVA谱特征值的QTL分析
Table 4 QTL detected for RVA in CSSLs population.
位点 QTL | 区间 Marker interval | LOD值LOD value | 贡献率PVE/% | ||||||
---|---|---|---|---|---|---|---|---|---|
E1 | E2 | E3 | E4 | E1 | E2 | E3 | E4 | ||
qPKV1 | RM6292-RM5362 | 3.08 | 10.44 | ||||||
qPKV3 | RM6297-RM1278 | 6.30 | 40.98 | ||||||
qPKV6.1 | RM1369-RM510 | 3.42 | 25.32 | ||||||
qPKV6.2 | RM510-RM1163 | 5.61 | 20.35 | ||||||
qPKV10 | RM6737-RM25923 | 4.19 | 22.55 | ||||||
qPKV11 | STS-OSJNBa0095P22-RM5766 | 3.05 | 15.69 | ||||||
qHPV3 | RM6297-RM1278 | 5.61 | 17.01 | ||||||
qHPV6 | RM1369-RM510 | 12.22 | 4.98 | 12.64 | 30.11 | 64.75 | 30.21 | 53.41 | 76.71 |
qHPV8 | RM6838-RM5767 | 2.79 | 3.21 | ||||||
qBDV3 | RM15887-RM15903 | 3.79 | 3.34 | ||||||
qBDV5 | RM3345-RM17954 | 3.72 | 14.25 | ||||||
qBDV6 | RM1369-RM510 | 25.94 | 4.98 | 11.39 | 35.00 | 89.05 | 32.21 | 61.47 | 77.94 |
qCPV3.1 | RM15887-RM15903 | 2.78 | 4.40 | ||||||
qCPV3.2 | RM6297-RM1278 | 11.11 | 13.98 | ||||||
qCPV6 | RM1369-RM510 | 20.80 | 10.46 | 22.26 | 45.45 | 80.40 | 50.81 | 60.34 | 90.23 |
qCPV8 | RM5556-RM6838 | 11.58 | 14.92 | ||||||
qSBV2 | RM5529-RM7581 | 2.88 | 5.12 | ||||||
qSBV3 | RM15887-RM15903 | 5.06 | 1.61 | ||||||
qSBV5 | RM3345-RM17954 | 3.54 | 6.45 | ||||||
qSBV6.1 | RM1369-RM510 | 33.68 | 19.23 | 22.89 | 56.05 | 94.34 | 70.67 | 85.28 | 90.49 |
qSBV6.2 | RM510-RM1163 | 5.81 | 1.89 | ||||||
qSBV8 | RM7049-RM7556 | 3.37 | 1.03 | ||||||
qCSV3 | RM15887-RM15903 | 3.23 | 3.72 | ||||||
qCSV5 | RM3476-RM3790 | 2.74 | 2.97 | ||||||
qCSV6 | RM1369-RM510 | 24.77 | 12.92 | 11.12 | 30.99 | 85.33 | 63.53 | 60.61 | 76.79 |
qPaT4 | RM3839-RM303 | 3.53 | 5.04 | ||||||
qPaT6.1 | RM1369-RM510 | 2.70 | 11.27 | 15.05 | 19.62 | ||||
qPaT6.2 | RM510-RM1163 | 4.63 | 32.63 | ||||||
qPaT10 | RM6737-RM25923 | 4.59 | 30.13 | ||||||
qPaT12 | RM3331-RM5479 | 3.90 | 20.99 | 22.88 | 48.48 | ||||
qPeT6 | RM1369-RM510 | 11.05 | 3.59 | 48.48 | 16.77 | ||||
qPeT7.1 | RM1132-RM8261 | 3.51 | 10.78 | ||||||
qPeT7.2 | STS-OJ1123_C12-RM5752 | 4.47 | 23.25 | ||||||
qPeT7.3 | RM7273-RM3404 | 2.95 | 14.44 |
图2 稻米品质性状QTL对应片段代换系与背景亲本昌恢121的表型差异比较*和**分别表示昌恢121的品质性状与目标片段代换系之间的差异达5%和1%显著水平。NC和HN分别表示江西南昌和海南。
Fig. 2. Differences of phenotypic values of rice quality traits between genetic background parent Changhui 121 and the CSSLs harboring the QTL alleles. *and** mean significant difference at 5% and 1% levels between rice quality traits of Changhui 121 and the CSSLs harboring the QTL alleles. NC and HN mean Nanchang City, Jiangxi Province and Hainan Province, respectively.
染色体 Chr | 主效QTL簇 Major QTL cluster | 区间 Marker interval | 性状 Trait | 多效性QTL位点 Pleiotropic QTL cluster |
---|---|---|---|---|
2 | qPGWC2 | RM5529–RM7581 | PGWC, SBV | qPGWC2, qSBV2 |
3 | qCPV3.2 | RM6297–RM1278 | CPV, PKV, HPV | qPKV3, qHPV3, qCPV3.2 |
3 | qSBV3 | RM15887–RM15903 | BDV, CPV, SBV, CSV | qBDV3, qCPV3.1, qSBV3, qCSV3 |
5 | qBDV5 | RM3345–RM17954 | BDV, SBV, PGWC, GC | qPGWC5, qGC5, qBDV5, qSBV5 |
6 | qSBV6.1 | RM1369–RM510 | GC, PKV, HPV, BDV, CPV, SBV, CSV, PaT, PeT | qGC6.1, qPKV6.1, qHPV6, qBDV6, qCPV6, qSBV6.1, qCSV6, qPaT6.1, qPeT6 |
6 | qSBV6.2 | RM510–RM1163 | GC, PKV, SBV, PaT | qGC6.2, qPKV6.2, qSBV6.2, qPaT6.2 |
10 | qPaT10 | RM6737–RM25923 | PKV, PaT, PC | qPKV10, qPaT10, qPC10.1 |
表5 水稻品质多效性区域分析
Table 5 Grain quality traits QTL hotspot and pleiotropic region in rice.
染色体 Chr | 主效QTL簇 Major QTL cluster | 区间 Marker interval | 性状 Trait | 多效性QTL位点 Pleiotropic QTL cluster |
---|---|---|---|---|
2 | qPGWC2 | RM5529–RM7581 | PGWC, SBV | qPGWC2, qSBV2 |
3 | qCPV3.2 | RM6297–RM1278 | CPV, PKV, HPV | qPKV3, qHPV3, qCPV3.2 |
3 | qSBV3 | RM15887–RM15903 | BDV, CPV, SBV, CSV | qBDV3, qCPV3.1, qSBV3, qCSV3 |
5 | qBDV5 | RM3345–RM17954 | BDV, SBV, PGWC, GC | qPGWC5, qGC5, qBDV5, qSBV5 |
6 | qSBV6.1 | RM1369–RM510 | GC, PKV, HPV, BDV, CPV, SBV, CSV, PaT, PeT | qGC6.1, qPKV6.1, qHPV6, qBDV6, qCPV6, qSBV6.1, qCSV6, qPaT6.1, qPeT6 |
6 | qSBV6.2 | RM510–RM1163 | GC, PKV, SBV, PaT | qGC6.2, qPKV6.2, qSBV6.2, qPaT6.2 |
10 | qPaT10 | RM6737–RM25923 | PKV, PaT, PC | qPKV10, qPaT10, qPC10.1 |
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