中国水稻科学 ›› 2022, Vol. 36 ›› Issue (3): 259-268.DOI: 10.16819/j.1001-7216.2022
刘进1,2, 崔迪2, 余丽琴1, 张立娜2, 周慧颖1, 马小定2, 胡佳晓1, 韩冰2, 韩龙植2,*(), 黎毛毛1,*()
收稿日期:
2021-07-29
修回日期:
2021-08-19
出版日期:
2022-05-10
发布日期:
2022-05-11
通讯作者:
韩龙植,黎毛毛
基金资助:
LIU Jin1,2, CUI Di2, YU Liqin1, ZHANG Lina2, ZHOU Huiying1, MA Xiaoding2, HU Jiaxiao1, HAN Bing2, HAN Longzhi2,*(), LI Maomao1,*()
Received:
2021-07-29
Revised:
2021-08-19
Online:
2022-05-10
Published:
2022-05-11
Contact:
HAN Longzhi, LI Maomao
摘要:
【目的】鉴定和筛选水稻极端耐热种质或基因,为培育耐高温水稻新品种提供技术支撑。【方法】以耐热等级和幼苗存活率为指标对不同类型水稻苗期耐热性进行鉴定评价,以筛选和鉴定耐热种质资源及主效QTL。【结果】不同类型水稻品种苗期耐热性存在明显差异,籼稻品种耐热性明显强于粳稻品种,籼稻和粳稻品种均存在极端耐热和极端敏感种质资源;共筛选出嘉育253、中优早8号、秀水09等20份耐热种质资源,高温处理后幼苗存活率和生长发育基本不受影响。RIL群体双亲耐热等级和幼苗存活率存在极显著差异,中优早8号耐热性较强,植株基本无枯死,龙稻5号对高温胁迫较敏感,不同株系间苗期耐热性存在较大幅度变异;共检测到12个苗期耐热相关QTL,分布于第1、3、4、5和8染色体上,耐热等级和存活率QTL存在明显的遗传重叠,主效QTL簇qHTS4和qHTS8表型贡献率较大。基于QTL初步定位结果,利用相对剩余杂合体RHL-F2群体,在第8染色体RM5808-RM556标记区域鉴定了一个苗期耐热性主效QTL qHTS8,该区域对苗期耐热性具有较强调控效应。【结论】筛选出20份苗期耐热性较强的水稻种质资源,鉴定了12个苗期耐热相关的QTL,定位和验证了一个调控水稻苗期耐热性的主效QTL qHTS8,研究结果可为水稻苗期耐热性生理生化机理与分子遗传机制的研究及育种利用奠定基础。
刘进, 崔迪, 余丽琴, 张立娜, 周慧颖, 马小定, 胡佳晓, 韩冰, 韩龙植, 黎毛毛. 水稻苗期耐热种质资源筛选及QTL定位[J]. 中国水稻科学, 2022, 36(3): 259-268.
LIU Jin, CUI Di, YU Liqin, ZHANG Lina, ZHOU Huiying, MA Xiaoding, HU Jiaxiao, HAN Bing, HAN Longzhi, LI Maomao. Screening and QTL Mapping of Heat-tolerant Rice (Oryza sativa L.) Germplasm Resources at Seedling Stage[J]. Chinese Journal OF Rice Science, 2022, 36(3): 259-268.
图1 水稻苗期耐热鉴定 A~C分别为正常温度培养水稻苗期培养环境及植株长势;D~F分别为45℃处理恢复后表型、耐热等级和极端材料表型。
Fig. 1. Identification of heat tolerance at seedling stage in rice. A-C, Plant phenotype at seedling stage under control; D-F, Plant phenotype, heat tolerance grade and extreme material after exposure to 45℃.
性状 Trait | 亚群 Group | 品种数量 Number of varieties | 均值±标准差 Mean±SD | 变幅 Range | 峰度 Skewness | 偏度 Kurtosis |
---|---|---|---|---|---|---|
耐热等级 Heat tolerance grade | 总体 Total | 738 | 6.04±2.38 | 1~9 | -0.498 | -0.749 |
籼稻 indica | 387 | 5.40±2.73 | 1~9 | -0.111 | -1.298 | |
粳稻 japonica | 351 | 6.74±1.67** | 3~9 | -0.417 | -0.568 | |
存活率 Survival rate under heat stress | 总体 Total | 738 | 42.41±30.42 | 0~100 | 0.226 | -1.174 |
籼稻 indica | 387 | 50.35±31.97 | 0~100 | -0.090 | -1.343 | |
粳稻 japonica | 351 | 33.64±25.98** | 0~100 | 0.436 | -0.787 |
表1 水稻种质资源苗期耐热性鉴定
Table 1. Screening of heat-tolerant rice germplasm resources at seedling stage.
性状 Trait | 亚群 Group | 品种数量 Number of varieties | 均值±标准差 Mean±SD | 变幅 Range | 峰度 Skewness | 偏度 Kurtosis |
---|---|---|---|---|---|---|
耐热等级 Heat tolerance grade | 总体 Total | 738 | 6.04±2.38 | 1~9 | -0.498 | -0.749 |
籼稻 indica | 387 | 5.40±2.73 | 1~9 | -0.111 | -1.298 | |
粳稻 japonica | 351 | 6.74±1.67** | 3~9 | -0.417 | -0.568 | |
存活率 Survival rate under heat stress | 总体 Total | 738 | 42.41±30.42 | 0~100 | 0.226 | -1.174 |
籼稻 indica | 387 | 50.35±31.97 | 0~100 | -0.090 | -1.343 | |
粳稻 japonica | 351 | 33.64±25.98** | 0~100 | 0.436 | -0.787 |
编号Number | 品种名称 Cultivar | 耐热等级 HTG | HTSR /% | 编号 Number | 品种名称Cultivar | 耐热等级 HTG | HTSR /% | ||
---|---|---|---|---|---|---|---|---|---|
L1 | 嘉育253 Jiayu 253 | 1 | 100.0 | L11 | 秀水9号 Xiushui 9 | 3 | 93.8 | ||
L2 | 早恢6号 Zaohui 6 | 1 | 100.0 | L12 | 苏资16 Suzi 16 | 4 | 87.5 | ||
L3 | 龙红3号 Longhong 3 | 1 | 100.0 | L13 | 秀水12 Xiushui 12 | 4 | 87.5 | ||
L4 | 闽泉2号 Minquan 2 | 1 | 100.0 | L14 | 云粳38 Yunjing 38 | 3 | 87.5 | ||
L5 | 湘晚籼11 Xiangwanxian 11 | 1 | 100.0 | L15 | 秀水48 Xiushui 48 | 4 | 75.0 | ||
L6 | 中优早8号 Zhongyouzao 8 | 1 | 97.0 | L16 | 嘉花1号 Jiahua 1 | 4 | 75.0 | ||
L7 | 梅峰23号 Meifeng 23 | 1 | 90.6 | L17 | 南粳46 Nanjing 46 | 3 | 73.3 | ||
L8 | 龙菲313 Longfei 313 | 1 | 90.6 | L18 | 甬糯34 Yongnuo 34 | 4 | 68.8 | ||
L9 | 莲塘早 Liantangzao | 3 | 86.9 | L19 | 盐恢559 Yanhui 559 | 4 | 64.3 | ||
L10 | 赣早籼58 Ganzaoxian 58 | 3 | 83.3 | L20 | 庄育3号 Zhuangyu 3 | 4 | 60.0 |
表2 水稻苗期极端耐热的种质资源
Table 2. Extreme heat resistant germplasm resources at seedling stage in rice.
编号Number | 品种名称 Cultivar | 耐热等级 HTG | HTSR /% | 编号 Number | 品种名称Cultivar | 耐热等级 HTG | HTSR /% | ||
---|---|---|---|---|---|---|---|---|---|
L1 | 嘉育253 Jiayu 253 | 1 | 100.0 | L11 | 秀水9号 Xiushui 9 | 3 | 93.8 | ||
L2 | 早恢6号 Zaohui 6 | 1 | 100.0 | L12 | 苏资16 Suzi 16 | 4 | 87.5 | ||
L3 | 龙红3号 Longhong 3 | 1 | 100.0 | L13 | 秀水12 Xiushui 12 | 4 | 87.5 | ||
L4 | 闽泉2号 Minquan 2 | 1 | 100.0 | L14 | 云粳38 Yunjing 38 | 3 | 87.5 | ||
L5 | 湘晚籼11 Xiangwanxian 11 | 1 | 100.0 | L15 | 秀水48 Xiushui 48 | 4 | 75.0 | ||
L6 | 中优早8号 Zhongyouzao 8 | 1 | 97.0 | L16 | 嘉花1号 Jiahua 1 | 4 | 75.0 | ||
L7 | 梅峰23号 Meifeng 23 | 1 | 90.6 | L17 | 南粳46 Nanjing 46 | 3 | 73.3 | ||
L8 | 龙菲313 Longfei 313 | 1 | 90.6 | L18 | 甬糯34 Yongnuo 34 | 4 | 68.8 | ||
L9 | 莲塘早 Liantangzao | 3 | 86.9 | L19 | 盐恢559 Yanhui 559 | 4 | 64.3 | ||
L10 | 赣早籼58 Ganzaoxian 58 | 3 | 83.3 | L20 | 庄育3号 Zhuangyu 3 | 4 | 60.0 |
性状 Trait | 亲本 Parent | 重组自交系群体 RIL population | |||||
---|---|---|---|---|---|---|---|
龙稻5号 Longdao 5 | 中优早8号 Zhongyouzao 8 | 均值±标准差 Mean±SD | 变幅 Range | 峰度 Skewness | 偏度 Kurtosis | ||
耐热等级HTG | 8.0±1.2 | 1.7±1.0** | 6.5±1.8 | 1.7~9.0 | -0.55 | -0.28 | |
HTSR/% | 17.5±6.5 | 90.8±3.0** | 39.1±16.5 | 0.0~100.0 | 0.23 | -0.77 |
表3 亲本和RIL群体苗期耐热性表型分析
Table 3. Phenotype of heat tolerance from the parents and RIL populations at seedling stage.
性状 Trait | 亲本 Parent | 重组自交系群体 RIL population | |||||
---|---|---|---|---|---|---|---|
龙稻5号 Longdao 5 | 中优早8号 Zhongyouzao 8 | 均值±标准差 Mean±SD | 变幅 Range | 峰度 Skewness | 偏度 Kurtosis | ||
耐热等级HTG | 8.0±1.2 | 1.7±1.0** | 6.5±1.8 | 1.7~9.0 | -0.55 | -0.28 | |
HTSR/% | 17.5±6.5 | 90.8±3.0** | 39.1±16.5 | 0.0~100.0 | 0.23 | -0.77 |
图3 水稻RIL群体苗期耐热性表型鉴定及相关性分析 A~C为高温处理后表型性状; D-幼苗耐热等级; E-幼苗存活率; F―耐热等级和幼苗存活率相关性。
Fig. 3. Identification of heat tolerance of RIL population and correlation at seedling stage. A-C is the phenotypic after high temperature treatment; D, Heat tolerance grade; E, Seedling survival distribution; F, Correlation between heat tolerance grade and seedling survival rate under heat stress.
性状 Trait | 位点 Locus | 位置 Position | 标记 Marker | LOD值 LOD value | 贡献率 Percentage variance explained / % | 加性效应 Additive effect |
---|---|---|---|---|---|---|
耐热等级 Heat tolerance grade | qHTG1.1 1 | 50.5 | RI02519-RM580 | 2.56 | 4.11 | 0.40 |
qHTG1.2 | 152.5 | RM1198-RM1361.1 | 2.51 | 4.76 | 0.53 | |
qHTG4 | 123.1 | RM3648-RM1113 | 2.54 | 12.46 | -0.41 | |
qHTG5.1 | 111.0 | STS5.3-RM26 | 2.79 | 3.25 | -0.47 | |
qHTG5.2 | 132.0 | RM19221-RM334 | 2.50 | 3.54 | -0.49 | |
qHTG8 | 81.5 | RM7285-RM6976 | 4.34 | 14.57 | -1.63 | |
幼苗存活率 Survival rate under heat stress | qHTSR1 | 50.5 | RI02519-RM580 | 2.69 | 4.69 | -6.73 |
qHTSR3 | 87.5 | MM3641-MM3720 | 2.54 | 2.39 | 6.20 | |
qHTSR4.1 | 95.1 | R4M50-STS4.3 | 2.61 | 3.50 | 5.69 | |
qHTSR4.2 | 123.1 | RM3648-RM1113 | 2.79 | 13.09 | 7.06 | |
qHTSR5 | 108.0 | RM3321-RM3616 | 2.56 | 3.23 | 7.21 | |
qHTSR8 | 80.5 | RM7285-RM6976 | 5.05 | 15.30 | 19.69 |
表4 水稻苗期耐热性QTL分析
Table 4. QTL for heat tolerance were detected at seedling stage.
性状 Trait | 位点 Locus | 位置 Position | 标记 Marker | LOD值 LOD value | 贡献率 Percentage variance explained / % | 加性效应 Additive effect |
---|---|---|---|---|---|---|
耐热等级 Heat tolerance grade | qHTG1.1 1 | 50.5 | RI02519-RM580 | 2.56 | 4.11 | 0.40 |
qHTG1.2 | 152.5 | RM1198-RM1361.1 | 2.51 | 4.76 | 0.53 | |
qHTG4 | 123.1 | RM3648-RM1113 | 2.54 | 12.46 | -0.41 | |
qHTG5.1 | 111.0 | STS5.3-RM26 | 2.79 | 3.25 | -0.47 | |
qHTG5.2 | 132.0 | RM19221-RM334 | 2.50 | 3.54 | -0.49 | |
qHTG8 | 81.5 | RM7285-RM6976 | 4.34 | 14.57 | -1.63 | |
幼苗存活率 Survival rate under heat stress | qHTSR1 | 50.5 | RI02519-RM580 | 2.69 | 4.69 | -6.73 |
qHTSR3 | 87.5 | MM3641-MM3720 | 2.54 | 2.39 | 6.20 | |
qHTSR4.1 | 95.1 | R4M50-STS4.3 | 2.61 | 3.50 | 5.69 | |
qHTSR4.2 | 123.1 | RM3648-RM1113 | 2.79 | 13.09 | 7.06 | |
qHTSR5 | 108.0 | RM3321-RM3616 | 2.56 | 3.23 | 7.21 | |
qHTSR8 | 80.5 | RM7285-RM6976 | 5.05 | 15.30 | 19.69 |
图4 水稻耐热等级(黑色)和幼苗存活率(红色)QTL在RIL群体中的染色体分布
Fig. 4. Location of QTL for heat tolerance grade(black) and seedling survival rate(red) detected at seedling stage on RIL populations genetic map.
图5 水稻苗期耐热主效qHTS8定位相关的剩余杂合体基因型
Fig. 5. Residual heterozygous genotypes for identification of major QTL qHTS8 for heat tolerance at seedling stage in rice.
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