中国水稻科学 ›› 2022, Vol. 36 ›› Issue (2): 159-170.DOI: 10.16819/j.1001-7216.2021.210204
黄涛1,2, 王燕宁1, 钟奇1, 程琴1, 杨朦朦1, 王鹏1, 吴光亮1, 黄诗颖1, 李才敬1, 余剑峰1, 贺浩华1,3,*(), 边建民1,3,*(
)
收稿日期:
2021-02-06
修回日期:
2021-05-02
出版日期:
2022-03-10
发布日期:
2022-03-11
通讯作者:
贺浩华,边建民
基金资助:
HUANG Tao1,2, WANG Yanning1, ZHONG Qi1, CHENG Qin1, YANG Mengmeng1, WANG Peng1, WU Guangliang1, HUANG Shiying1, LI Caijing1, YU Jianfeng1, HE Haohua1,3,*(), BIAN Jianmin1,3,*(
)
Received:
2021-02-06
Revised:
2021-05-02
Online:
2022-03-10
Published:
2022-03-11
Contact:
HE Haohua, BIAN Jianmin
摘要:
【目的】 挖掘水稻粒重和粒型相关性状QTL,对于解析水稻籽粒遗传机理具有重要作用。【方法】 本研究以籼稻9311为受体、粳稻日本晴为供体构建的染色体片段置换系(Chromosome Segment Substitution Lines, CSSLs)群体为材料,在4个环境下对控制稻谷与糙米的粒重和粒型QTL进行了定位分析。【结果】 共检测到77个控制水稻粒重和粒型的QTL,贡献率为4.62%~51.01%,其中19个QTL的增效等位基因来自日本晴,58个QTL的增效等位基因来自9311。这些QTL分布在水稻10条染色体的46个区域,其中16个区域为多效性位点。在两个及两个以上环境中重复检测到的QTL有14个,其中qGW5.1和qLW5能够在4个环境中稳定表达,且位于同一染色体区域;qBRL3.2和qGL4.1为新鉴定的影响水稻粒重和粒型的QTL。【结论】 本研究结果为后续克隆这些QTL和解析水稻粒重和粒型遗传机理奠定了基础。
黄涛, 王燕宁, 钟奇, 程琴, 杨朦朦, 王鹏, 吴光亮, 黄诗颖, 李才敬, 余剑峰, 贺浩华, 边建民. 利用染色体片段置换系群体定位和分析水稻粒重和粒型QTL[J]. 中国水稻科学, 2022, 36(2): 159-170.
HUANG Tao, WANG Yanning, ZHONG Qi, CHENG Qin, YANG Mengmeng, WANG Peng, WU Guangliang, HUANG Shiying, LI Caijing, YU Jianfeng, HE Haohua, BIAN Jianmin. Mapping and Analysis of QTLs for Rice Grain Weight and Grain Shape Using Chromosome Segment Substitution Line Population[J]. Chinese Journal OF Rice Science, 2022, 36(2): 159-170.
性状 Trait | 环境Environment | 亲本 Parent | 置换系群体 CSSL | |||||
---|---|---|---|---|---|---|---|---|
9311 | 日本晴 Nipponbare | 均值±标准Mean±SD | 变幅 Range | 峰度Kurtosis | 偏度Skewness | |||
稻谷粒长 | E1 | 9.86±0.47** | 7.38±1.56 | 9.59±0.35 | 8.37~10.69 | -0.46 | 2.58 | |
Grain length / mm | E2 | 9.76±0.08** | 7.68±0.16 | 9.31±0.28 | 8.23~9.77 | -1.66 | 3.99 | |
E3 | 9.79±0.16** | 7.67±0.19 | 9.52±0.33 | 8.05~10.14 | -1.79 | 5.19 | ||
E4 | 9.80±0.24** | 7.63±0.29 | 9.78±0.31 | 8.71~10.30 | -1.06 | 1.42 | ||
稻谷粒宽 | E1 | 2.79±0.12* | 3.24±0.60 | 2.78±0.13 | 2.55~3.53 | 2.33 | 10.17 | |
Grain width / mm | E2 | 2.48±0.12* | 3.30±0.56 | 2.99±0.12 | 2.69~3.61 | 2.00 | 8.05 | |
E3 | 2.59±0.19* | 3.17±0.00 | 2.81±0.13 | 2.43~3.38 | 1.83 | 6.11 | ||
E4 | 2.65±0.14* | 3.13±0.42 | 2.83±0.11 | 2.56~3.27 | 1.43 | 4.11 | ||
稻谷长宽比 | E1 | 3.55±0.27** | 2.29±0.13 | 3.49±0.19 | 2.62~3.78 | -1.91 | 4.62 | |
Length-width ratio | E2 | 3.47±0.12** | 2.31±0.23 | 3.14±0.16 | 2.42~3.38 | 1.60 | 3.70 | |
E3 | 3.78±0.11** | 2.42±0.19 | 3.41±0.19 | 2.71~3.71 | -1.59 | 2.73 | ||
E4 | 3.70±0.05** | 2.34±0.14 | 3.47±0.18 | 2.60~3.80 | -1.82 | 5.04 | ||
稻谷粒厚 | E1 | 2.26±0.09 | 2.17±0.05 | 2.19±0.05 | 2.04~2.34 | 0.07 | 0.64 | |
Grain thickness / mm | E2 | 2.14±0.10 | 2.09±0.06 | 2.18±0.05 | 2.05~2.33 | 0.20 | 0.12 | |
E3 | 2.09±0.05 | 2.19±0.09 | 2.12±0.06 | 1.93~2.24 | -0.54 | 0.05 | ||
E4 | 1.99±0.06 | 2.01±0.12 | 2.10±0.05 | 1.95~2.20 | -0.33 | 0.05 | ||
稻谷千粒重 | E1 | 31.02±2.37** | 22.44±1.69 | 29.51±2.29 | 22.17~34.21 | -0.64 | 0.70 | |
1000-grain weight / g | E2 | 31.36±3.06** | 22.26±2.01 | 29.63±1.97 | 23.24~34.90 | -0.40 | 0.77 | |
E3 | 30.39±2.74** | 21.86±1.49 | 27.39±2.42 | 19.71~33.65 | -0.21 | 0.48 | ||
E4 | 32.53±2.36** | 22.65±1.84 | 29.46±2.16 | 23.46~33.49 | -0.58 | 0.41 | ||
糙米长 | E1 | 6.87±0.31** | 4.88±0.36 | 6.91±0.23 | 6.14~7.30 | -1.50 | 2.67 | |
Brown rice length | E2 | 6.84±0.41** | 0.86±0.49 | 6.84±0.24 | 5.94~7.36 | -1.64 | 3.57 | |
/mm | E3 | 6.91±0.46** | 4.99±0.61 | 6.93±0.24 | 5.96~7.39 | -1.54 | 3.65 | |
E4 | 6.91±0.37** | 4.70±0.23 | 6.92±0.27 | 5.97~7.54 | -0.57 | 0.55 | ||
糙米宽 | E1 | 2.38±0.12* | 2.70±0.08 | 2.36±0.11 | 2.17~2.92 | 2.04 | 7.85 | |
Brown rice width | E2 | 2.27±0.14* | 2.67±0.13 | 2.52±0.09 | 2.31~3.01 | 1.90 | 9.25 | |
/mm | E3 | 2.32±0.19* | 2.78±0.05 | 2.40±0.09 | 2.16~2.79 | 1.50 | 5.41 | |
E4 | 2.46±0.11* | 2.75±0.06 | 2.36±0.11 | 2.07~2.89 | 1.17 | 4.67 | ||
糙米长宽比 | E1 | 2.98±0.21** | 1.87±0.19 | 2.95±0.17 | 2.17~3.21 | -1.67 | 3.84 | |
Length-to-width ratio | E2 | 2.94±0.23** | 1.84±0.17 | 2.72±0.14 | 2.11~2.99 | -1.56 | 4.07 | |
of brown rice | E3 | 2.98±0.31** | 1.81±0.16 | 2.91±0.15 | 2.33~3.15 | -1.53 | 3.09 | |
E4 | 2.76±0.27** | 1.86±0.19 | 2.95±0.14 | 2.26~3.16 | -1.95 | 5.56 | ||
糙米厚 | E1 | 1.93±0.10 | 2.07±0.06* | 1.93±0.05 | 1.79~2.10 | 0.14 | 0.66 | |
Thickness of brown rice | E2 | 1.90±0.01 | 2.12±0.04* | 1.91±0.05 | 1.77~2.08 | 0.36 | 1.48 | |
/ mm | E3 | 1.85±0.07 | 1.96±0.06* | 1.87±0.05 | 1.73~2.07 | 0.10 | 2.63 | |
E4 | 1.91±0.06 | 2.10±0.05* | 1.88±0.05 | 1.77~2.06 | 0.37 | 0.38 | ||
糙米千粒重 | E1 | 23.55±0.49* | 21.88±0.32 | 24.64±1.57 | 20.47~29.46 | 0.03 | 0.33 | |
1000-grain weight | E2 | 22.86±0.09* | 21.76±0.36 | 24.32±1.37 | 20.34~28.20 | -0.21 | 0.72 | |
of brown rice / g | E3 | 22.29±0.14* | 20.35±0.60 | 23.14±1.61 | 18.19~27.60 | -0.10 | 0.31 | |
E4 | 23.67±0.52* | 21.66±0.54 | 2.83±1.56 | 19.63~26.95 | -0.39 | 0.00 |
表1 四个环境下亲本9311和日本晴及CSSL群体粒重和粒型表现
Table 1 Grain weight and shape performance of 9311 and Nipponbare and CSSLs population in four environments.
性状 Trait | 环境Environment | 亲本 Parent | 置换系群体 CSSL | |||||
---|---|---|---|---|---|---|---|---|
9311 | 日本晴 Nipponbare | 均值±标准Mean±SD | 变幅 Range | 峰度Kurtosis | 偏度Skewness | |||
稻谷粒长 | E1 | 9.86±0.47** | 7.38±1.56 | 9.59±0.35 | 8.37~10.69 | -0.46 | 2.58 | |
Grain length / mm | E2 | 9.76±0.08** | 7.68±0.16 | 9.31±0.28 | 8.23~9.77 | -1.66 | 3.99 | |
E3 | 9.79±0.16** | 7.67±0.19 | 9.52±0.33 | 8.05~10.14 | -1.79 | 5.19 | ||
E4 | 9.80±0.24** | 7.63±0.29 | 9.78±0.31 | 8.71~10.30 | -1.06 | 1.42 | ||
稻谷粒宽 | E1 | 2.79±0.12* | 3.24±0.60 | 2.78±0.13 | 2.55~3.53 | 2.33 | 10.17 | |
Grain width / mm | E2 | 2.48±0.12* | 3.30±0.56 | 2.99±0.12 | 2.69~3.61 | 2.00 | 8.05 | |
E3 | 2.59±0.19* | 3.17±0.00 | 2.81±0.13 | 2.43~3.38 | 1.83 | 6.11 | ||
E4 | 2.65±0.14* | 3.13±0.42 | 2.83±0.11 | 2.56~3.27 | 1.43 | 4.11 | ||
稻谷长宽比 | E1 | 3.55±0.27** | 2.29±0.13 | 3.49±0.19 | 2.62~3.78 | -1.91 | 4.62 | |
Length-width ratio | E2 | 3.47±0.12** | 2.31±0.23 | 3.14±0.16 | 2.42~3.38 | 1.60 | 3.70 | |
E3 | 3.78±0.11** | 2.42±0.19 | 3.41±0.19 | 2.71~3.71 | -1.59 | 2.73 | ||
E4 | 3.70±0.05** | 2.34±0.14 | 3.47±0.18 | 2.60~3.80 | -1.82 | 5.04 | ||
稻谷粒厚 | E1 | 2.26±0.09 | 2.17±0.05 | 2.19±0.05 | 2.04~2.34 | 0.07 | 0.64 | |
Grain thickness / mm | E2 | 2.14±0.10 | 2.09±0.06 | 2.18±0.05 | 2.05~2.33 | 0.20 | 0.12 | |
E3 | 2.09±0.05 | 2.19±0.09 | 2.12±0.06 | 1.93~2.24 | -0.54 | 0.05 | ||
E4 | 1.99±0.06 | 2.01±0.12 | 2.10±0.05 | 1.95~2.20 | -0.33 | 0.05 | ||
稻谷千粒重 | E1 | 31.02±2.37** | 22.44±1.69 | 29.51±2.29 | 22.17~34.21 | -0.64 | 0.70 | |
1000-grain weight / g | E2 | 31.36±3.06** | 22.26±2.01 | 29.63±1.97 | 23.24~34.90 | -0.40 | 0.77 | |
E3 | 30.39±2.74** | 21.86±1.49 | 27.39±2.42 | 19.71~33.65 | -0.21 | 0.48 | ||
E4 | 32.53±2.36** | 22.65±1.84 | 29.46±2.16 | 23.46~33.49 | -0.58 | 0.41 | ||
糙米长 | E1 | 6.87±0.31** | 4.88±0.36 | 6.91±0.23 | 6.14~7.30 | -1.50 | 2.67 | |
Brown rice length | E2 | 6.84±0.41** | 0.86±0.49 | 6.84±0.24 | 5.94~7.36 | -1.64 | 3.57 | |
/mm | E3 | 6.91±0.46** | 4.99±0.61 | 6.93±0.24 | 5.96~7.39 | -1.54 | 3.65 | |
E4 | 6.91±0.37** | 4.70±0.23 | 6.92±0.27 | 5.97~7.54 | -0.57 | 0.55 | ||
糙米宽 | E1 | 2.38±0.12* | 2.70±0.08 | 2.36±0.11 | 2.17~2.92 | 2.04 | 7.85 | |
Brown rice width | E2 | 2.27±0.14* | 2.67±0.13 | 2.52±0.09 | 2.31~3.01 | 1.90 | 9.25 | |
/mm | E3 | 2.32±0.19* | 2.78±0.05 | 2.40±0.09 | 2.16~2.79 | 1.50 | 5.41 | |
E4 | 2.46±0.11* | 2.75±0.06 | 2.36±0.11 | 2.07~2.89 | 1.17 | 4.67 | ||
糙米长宽比 | E1 | 2.98±0.21** | 1.87±0.19 | 2.95±0.17 | 2.17~3.21 | -1.67 | 3.84 | |
Length-to-width ratio | E2 | 2.94±0.23** | 1.84±0.17 | 2.72±0.14 | 2.11~2.99 | -1.56 | 4.07 | |
of brown rice | E3 | 2.98±0.31** | 1.81±0.16 | 2.91±0.15 | 2.33~3.15 | -1.53 | 3.09 | |
E4 | 2.76±0.27** | 1.86±0.19 | 2.95±0.14 | 2.26~3.16 | -1.95 | 5.56 | ||
糙米厚 | E1 | 1.93±0.10 | 2.07±0.06* | 1.93±0.05 | 1.79~2.10 | 0.14 | 0.66 | |
Thickness of brown rice | E2 | 1.90±0.01 | 2.12±0.04* | 1.91±0.05 | 1.77~2.08 | 0.36 | 1.48 | |
/ mm | E3 | 1.85±0.07 | 1.96±0.06* | 1.87±0.05 | 1.73~2.07 | 0.10 | 2.63 | |
E4 | 1.91±0.06 | 2.10±0.05* | 1.88±0.05 | 1.77~2.06 | 0.37 | 0.38 | ||
糙米千粒重 | E1 | 23.55±0.49* | 21.88±0.32 | 24.64±1.57 | 20.47~29.46 | 0.03 | 0.33 | |
1000-grain weight | E2 | 22.86±0.09* | 21.76±0.36 | 24.32±1.37 | 20.34~28.20 | -0.21 | 0.72 | |
of brown rice / g | E3 | 22.29±0.14* | 20.35±0.60 | 23.14±1.61 | 18.19~27.60 | -0.10 | 0.31 | |
E4 | 23.67±0.52* | 21.66±0.54 | 2.83±1.56 | 19.63~26.95 | -0.39 | 0.00 |
图1 9311/日本晴CSSLs群体中稻谷和糙米粒型和粒重相关性状分布
Fig. 1. Frequency distribution of grain shape and grain weight related traits of 9311/Nipponbare CSSLs population.
图2 四个不同环境下CSSL群体稻谷和糙米的粒重和粒型相关性状间的相关性分析 颜色越红,表明相关性系数r越接近1,呈正相关;反之颜色越蓝,表明r越接近-1,呈负相关。GL-粒长;GW-粒宽;GT-粒厚;LW-长宽比;TGW-千粒重;BRL-糙米长;BRW-糙米宽;BRLW-糙米长宽比;BRTW-糙米千粒重。
Fig. 2. Correlation analysis between grain weight and shape related traits in CSSLs population under four environments. The redder the color, the closer the correlation coefficient r is to 1, meaning positive correlation. On the contrary, the bluer the color, the closer the correlation coefficient r is to -1, the negative correlation. GL, Grain length; GW, Grain width; GT, Grain thickness; LW, Length to width ratio; TGW, 1000-grain weight; BRL, Brown rice length; BRW, Brown rice width; BRLW, Length to width ratio of brown rice; BRTW, 1000-grain weight of brown rice.
QTL | 区间 Interval | E1 | E2 | E3 | E4 | ||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | ||||||||||||||||
qGL2 | M123-M124 | 3.32 | -0.29 | 7.63 | |||||||||||||||||||||||
qGL3.1 | M204-M205 | 6.61 | -0.48 | 17.64 | |||||||||||||||||||||||
qGL3.2 | M217-M218 | 3.40 | -0.30 | 11.16 | |||||||||||||||||||||||
qGL3.3 | 6.90 | -0.43 | 17.02 | ||||||||||||||||||||||||
qGL4.1 | M245-M246 | 3.01 | -0.38 | 7.48 | 3.04 | -0.34 | 6.94 | ||||||||||||||||||||
qGL4.2 | M255-M256 | 3.90 | -0.29 | 14.13 | |||||||||||||||||||||||
qGW1 | M9-M10 | 3.25 | -0.07 | 5.10 | |||||||||||||||||||||||
qGW2.1 | M142-M143 | 10.44 | 0.10 | 15.05 | |||||||||||||||||||||||
qGW2.2 | M157-M158 | 4.56 | 0.13 | 9.17 | |||||||||||||||||||||||
qGW5.2 | M334-M335 | 8.27 | 0.10 | 18.13 | |||||||||||||||||||||||
qGW5.1 | M328-M329 | 13.14 | 0.14 | 35.23 | 21.04 | 0.16 | 38.10 | 12.77 | 0.12 | 30.29 | 22.19 | 0.20 | 51.01 | ||||||||||||||
qGW7 | M491-M492 | 8.05 | 0.11 | 19.41 | |||||||||||||||||||||||
qGW8 | M537-M538 | 4.64 | -0.10 | 9.28 | |||||||||||||||||||||||
qGW9 | M553-M554 | 8.72 | -0.11 | 12.15 | |||||||||||||||||||||||
qGW10 | M639-M640 | 3.89 | 0.10 | 6.24 | |||||||||||||||||||||||
qLW2.1 | M136-M137 | 4.28 | -0.11 | 9.86 | |||||||||||||||||||||||
qLW2.2 | M138-M139 | 5.24 | -0.10 | 9.74 | |||||||||||||||||||||||
qLW3.2 | M204-M205 | 3.47 | -0.15 | 7.86 | |||||||||||||||||||||||
qLW3.1 | M217-M218 | 10.41 | -0.23 | 21.53 | |||||||||||||||||||||||
qLW3.3 | M220-M221 | 6.21 | -0.21 | 16.81 | |||||||||||||||||||||||
qLW4.1 | M245-M246 | 3.04 | -0.16 | 5.40 | |||||||||||||||||||||||
qLW4.2 | M255-M256 | 4.31 | -0.14 | 13.34 | |||||||||||||||||||||||
qLW4.3 | M276-M277 | 3.14 | -0.10 | 7.07 | |||||||||||||||||||||||
qLW5 | M328-M329 | 10.89 | -0.18 | 22.75 | 6.25 | -0.14 | 20.13 | 8.54 | -0.17 | 21.5 | 8.92 | -0.17 | 25.51 | ||||||||||||||
qLW7 | M491-M492 | 4.11 | -0.12 | 10.67 | |||||||||||||||||||||||
qLW10.0 | M643-M644 | 3.58 | -0.12 | 6.44 | 4.42 | -0.15 | 10.21 | ||||||||||||||||||||
qGT1.1 | M89-M90 | 6.42 | -0.05 | 14.72 | |||||||||||||||||||||||
qGT1.2 | M90-M91 | 3.52 | -0.04 | 10.40 | |||||||||||||||||||||||
qGT4 | M260-M261 | 3.08 | -0.04 | 6.59 | |||||||||||||||||||||||
qGT7.1 | M488-M489 | 3.18 | 0.06 | 10.01 | |||||||||||||||||||||||
qGT7.2 | M483-M484 | 3.31 | 0.05 | 12.42 | |||||||||||||||||||||||
qGT8.1 | M540-M541 | 3.23 | -0.04 | 10.17 | |||||||||||||||||||||||
qGT8.2 | M537-M538 | 7.02 | -0.09 | 16.30 | |||||||||||||||||||||||
qGT8.3 | M539-M540 | 4.89 | -0.04 | 15.12 | |||||||||||||||||||||||
qTGW2 | M153-M154 | 3.18 | 1.83 | 9.99 | |||||||||||||||||||||||
qTGW3 | M204-M205 | 4.90 | -2.66 | 12.58 | 3.11 | -2.16 | 9.67 | ||||||||||||||||||||
qTGW4.1 | M299-M300 | 3.29 | -1.51 | 7.96 | |||||||||||||||||||||||
qTGW4.2 | M308-M309 | 3.02 | -1.98 | 9.66 | |||||||||||||||||||||||
qTGW5.1 | M363-M364 | 13.76 | -10.14 | 11.55 | |||||||||||||||||||||||
qTGW5.2 | M374-M375 | 22.19 | 9.6 | 30.05 | |||||||||||||||||||||||
qTGW8.2 | M537-M538 | 3.29 | -2.38 | 10.36 | 4.00 | -2.47 | 12.68 | ||||||||||||||||||||
qTGW8.1 | M540-M541 | 4.47 | -1.69 | 11.37 | |||||||||||||||||||||||
qBRL1.1 | M61-M62 | 4.97 | -0.23 | 7.75 | |||||||||||||||||||||||
qBRL1.2 | M84-M85 | 3.08 | 0.13 | 4.62 | |||||||||||||||||||||||
qBRL2 | M123-M124 | 4.82 | -0.22 | 7.49 | |||||||||||||||||||||||
qBRL3.3 | M193-M194 | 5.86 | -0.18 | 9.30 | |||||||||||||||||||||||
qBRL3.1 | M204-M205 | 3.84 | -0.27 | 12.54 | |||||||||||||||||||||||
QTL | 区间 Interval | E1 | E2 | E3 | E4 | ||||||||||||||||||||||
LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | ||||||||||||||||
qBRL3.2 | M217-M218 | 12.22 | -0.38 | 30.96 | 9.17 | -0.28 | 15.59 | ||||||||||||||||||||
qBRL4.1 | M245-M246 | 3.94 | -0.28 | 8.40 | 4.70 | -0.27 | 7.29 | 4.02 | -0.36 | 10.74 | |||||||||||||||||
qBRL5.1 | M336-M337 | 4.18 | -0.20 | 11.41 | |||||||||||||||||||||||
qBRL8.1 | M537-M538 | 4.07 | -0.20 | 6.23 | |||||||||||||||||||||||
qBRL8.2 | M549-M550 | 3.66 | -0.24 | 9.72 | |||||||||||||||||||||||
qBRW2 | M143-M144 | 6.55 | 0.06 | 12.17 | |||||||||||||||||||||||
qBRW4 | M228-M229 | 3.44 | -0.08 | 10.98 | |||||||||||||||||||||||
qBRW5.2 | M328-M329 | 16.82 | 0.11 | 38.80 | |||||||||||||||||||||||
qBRW5.1 | M324-M325 | 5.62 | 0.10 | 19.85 | 15.31 | 0.12 | 43.56 | ||||||||||||||||||||
qBRW7 | M488-M489 | 4.60 | 0.10 | 10.46 | 5.16 | 0.16 | 17.04 | ||||||||||||||||||||
qBRW8.2 | M537-M538 | 4.39 | -0.07 | 7.82 | |||||||||||||||||||||||
qBRLW2.2 | M136-M137 | 3.31 | -0.08 | 10.38 | |||||||||||||||||||||||
qBRLW2.1 | M116-M117 | 3.33 | -0.08 | 6.32 | |||||||||||||||||||||||
qBRLW3.1 | M217-M218 | 12.09 | -0.19 | 27.46 | |||||||||||||||||||||||
qBRLW3.2 | M220-M221 | 6.22 | -0.18 | 20.65 | |||||||||||||||||||||||
qBRLW5 | M324-M325 | 3.22 | -0.12 | 11.89 | 9.64 | -0.14 | 20.80 | ||||||||||||||||||||
qBRLW7.1 | M489-M490 | 5.23 | -0.19 | 19.91 | |||||||||||||||||||||||
qBRLW7.2 | M488-M489 | 4.71 | -0.19 | 15.17 | |||||||||||||||||||||||
qBRT1 | M66-M67 | 5.70 | -0.05 | 12.94 | |||||||||||||||||||||||
qBRT2 | M170-M171 | 4.45 | -0.04 | 10.26 | |||||||||||||||||||||||
qBRT7.2 | M485-M486 | 5.09 | 0.05 | 11.41 | 4.33 | 0.06 | 13.69 | ||||||||||||||||||||
qBRT7.1 | M488-M489 | 4.59 | 0.07 | 16.39 | |||||||||||||||||||||||
qBRT8 | M544-M545 | 4.98 | -0.05 | 11.25 | |||||||||||||||||||||||
qBRTW1 | M66-M67 | 3.84 | -1.35 | 9.09 | |||||||||||||||||||||||
qBRTW2 | M153-M154 | 3.02 | 1.07 | 7.01 | |||||||||||||||||||||||
qBRTW3 | M204-M205 | 4.70 | -1.41 | 9.58 | 3.16 | -1.54 | 9.90 | ||||||||||||||||||||
qBRTW5.1 | M324-M325 | 6.81 | 1.24 | 14.48 | |||||||||||||||||||||||
qBRTW8.2 | M537-M538 | 5.36 | -1.86 | 13.03 | |||||||||||||||||||||||
qBRTW8.1 | M539-M540 | 4.27 | -1.17 | 8.79 | 4.77 | -1.38 | 15.64 | ||||||||||||||||||||
qBRTW11.1 | M652-M653 | 4.84 | -0.72 | 9.89 |
表2 利用CSSLs群体定位到的水稻粒型和粒重QTL
Table 2 QTL of rice grain shape and grain weight identified by CSSL population.
QTL | 区间 Interval | E1 | E2 | E3 | E4 | ||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | ||||||||||||||||
qGL2 | M123-M124 | 3.32 | -0.29 | 7.63 | |||||||||||||||||||||||
qGL3.1 | M204-M205 | 6.61 | -0.48 | 17.64 | |||||||||||||||||||||||
qGL3.2 | M217-M218 | 3.40 | -0.30 | 11.16 | |||||||||||||||||||||||
qGL3.3 | 6.90 | -0.43 | 17.02 | ||||||||||||||||||||||||
qGL4.1 | M245-M246 | 3.01 | -0.38 | 7.48 | 3.04 | -0.34 | 6.94 | ||||||||||||||||||||
qGL4.2 | M255-M256 | 3.90 | -0.29 | 14.13 | |||||||||||||||||||||||
qGW1 | M9-M10 | 3.25 | -0.07 | 5.10 | |||||||||||||||||||||||
qGW2.1 | M142-M143 | 10.44 | 0.10 | 15.05 | |||||||||||||||||||||||
qGW2.2 | M157-M158 | 4.56 | 0.13 | 9.17 | |||||||||||||||||||||||
qGW5.2 | M334-M335 | 8.27 | 0.10 | 18.13 | |||||||||||||||||||||||
qGW5.1 | M328-M329 | 13.14 | 0.14 | 35.23 | 21.04 | 0.16 | 38.10 | 12.77 | 0.12 | 30.29 | 22.19 | 0.20 | 51.01 | ||||||||||||||
qGW7 | M491-M492 | 8.05 | 0.11 | 19.41 | |||||||||||||||||||||||
qGW8 | M537-M538 | 4.64 | -0.10 | 9.28 | |||||||||||||||||||||||
qGW9 | M553-M554 | 8.72 | -0.11 | 12.15 | |||||||||||||||||||||||
qGW10 | M639-M640 | 3.89 | 0.10 | 6.24 | |||||||||||||||||||||||
qLW2.1 | M136-M137 | 4.28 | -0.11 | 9.86 | |||||||||||||||||||||||
qLW2.2 | M138-M139 | 5.24 | -0.10 | 9.74 | |||||||||||||||||||||||
qLW3.2 | M204-M205 | 3.47 | -0.15 | 7.86 | |||||||||||||||||||||||
qLW3.1 | M217-M218 | 10.41 | -0.23 | 21.53 | |||||||||||||||||||||||
qLW3.3 | M220-M221 | 6.21 | -0.21 | 16.81 | |||||||||||||||||||||||
qLW4.1 | M245-M246 | 3.04 | -0.16 | 5.40 | |||||||||||||||||||||||
qLW4.2 | M255-M256 | 4.31 | -0.14 | 13.34 | |||||||||||||||||||||||
qLW4.3 | M276-M277 | 3.14 | -0.10 | 7.07 | |||||||||||||||||||||||
qLW5 | M328-M329 | 10.89 | -0.18 | 22.75 | 6.25 | -0.14 | 20.13 | 8.54 | -0.17 | 21.5 | 8.92 | -0.17 | 25.51 | ||||||||||||||
qLW7 | M491-M492 | 4.11 | -0.12 | 10.67 | |||||||||||||||||||||||
qLW10.0 | M643-M644 | 3.58 | -0.12 | 6.44 | 4.42 | -0.15 | 10.21 | ||||||||||||||||||||
qGT1.1 | M89-M90 | 6.42 | -0.05 | 14.72 | |||||||||||||||||||||||
qGT1.2 | M90-M91 | 3.52 | -0.04 | 10.40 | |||||||||||||||||||||||
qGT4 | M260-M261 | 3.08 | -0.04 | 6.59 | |||||||||||||||||||||||
qGT7.1 | M488-M489 | 3.18 | 0.06 | 10.01 | |||||||||||||||||||||||
qGT7.2 | M483-M484 | 3.31 | 0.05 | 12.42 | |||||||||||||||||||||||
qGT8.1 | M540-M541 | 3.23 | -0.04 | 10.17 | |||||||||||||||||||||||
qGT8.2 | M537-M538 | 7.02 | -0.09 | 16.30 | |||||||||||||||||||||||
qGT8.3 | M539-M540 | 4.89 | -0.04 | 15.12 | |||||||||||||||||||||||
qTGW2 | M153-M154 | 3.18 | 1.83 | 9.99 | |||||||||||||||||||||||
qTGW3 | M204-M205 | 4.90 | -2.66 | 12.58 | 3.11 | -2.16 | 9.67 | ||||||||||||||||||||
qTGW4.1 | M299-M300 | 3.29 | -1.51 | 7.96 | |||||||||||||||||||||||
qTGW4.2 | M308-M309 | 3.02 | -1.98 | 9.66 | |||||||||||||||||||||||
qTGW5.1 | M363-M364 | 13.76 | -10.14 | 11.55 | |||||||||||||||||||||||
qTGW5.2 | M374-M375 | 22.19 | 9.6 | 30.05 | |||||||||||||||||||||||
qTGW8.2 | M537-M538 | 3.29 | -2.38 | 10.36 | 4.00 | -2.47 | 12.68 | ||||||||||||||||||||
qTGW8.1 | M540-M541 | 4.47 | -1.69 | 11.37 | |||||||||||||||||||||||
qBRL1.1 | M61-M62 | 4.97 | -0.23 | 7.75 | |||||||||||||||||||||||
qBRL1.2 | M84-M85 | 3.08 | 0.13 | 4.62 | |||||||||||||||||||||||
qBRL2 | M123-M124 | 4.82 | -0.22 | 7.49 | |||||||||||||||||||||||
qBRL3.3 | M193-M194 | 5.86 | -0.18 | 9.30 | |||||||||||||||||||||||
qBRL3.1 | M204-M205 | 3.84 | -0.27 | 12.54 | |||||||||||||||||||||||
QTL | 区间 Interval | E1 | E2 | E3 | E4 | ||||||||||||||||||||||
LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | LOD | 加性效应Additive | 贡献率PVE/% | ||||||||||||||||
qBRL3.2 | M217-M218 | 12.22 | -0.38 | 30.96 | 9.17 | -0.28 | 15.59 | ||||||||||||||||||||
qBRL4.1 | M245-M246 | 3.94 | -0.28 | 8.40 | 4.70 | -0.27 | 7.29 | 4.02 | -0.36 | 10.74 | |||||||||||||||||
qBRL5.1 | M336-M337 | 4.18 | -0.20 | 11.41 | |||||||||||||||||||||||
qBRL8.1 | M537-M538 | 4.07 | -0.20 | 6.23 | |||||||||||||||||||||||
qBRL8.2 | M549-M550 | 3.66 | -0.24 | 9.72 | |||||||||||||||||||||||
qBRW2 | M143-M144 | 6.55 | 0.06 | 12.17 | |||||||||||||||||||||||
qBRW4 | M228-M229 | 3.44 | -0.08 | 10.98 | |||||||||||||||||||||||
qBRW5.2 | M328-M329 | 16.82 | 0.11 | 38.80 | |||||||||||||||||||||||
qBRW5.1 | M324-M325 | 5.62 | 0.10 | 19.85 | 15.31 | 0.12 | 43.56 | ||||||||||||||||||||
qBRW7 | M488-M489 | 4.60 | 0.10 | 10.46 | 5.16 | 0.16 | 17.04 | ||||||||||||||||||||
qBRW8.2 | M537-M538 | 4.39 | -0.07 | 7.82 | |||||||||||||||||||||||
qBRLW2.2 | M136-M137 | 3.31 | -0.08 | 10.38 | |||||||||||||||||||||||
qBRLW2.1 | M116-M117 | 3.33 | -0.08 | 6.32 | |||||||||||||||||||||||
qBRLW3.1 | M217-M218 | 12.09 | -0.19 | 27.46 | |||||||||||||||||||||||
qBRLW3.2 | M220-M221 | 6.22 | -0.18 | 20.65 | |||||||||||||||||||||||
qBRLW5 | M324-M325 | 3.22 | -0.12 | 11.89 | 9.64 | -0.14 | 20.80 | ||||||||||||||||||||
qBRLW7.1 | M489-M490 | 5.23 | -0.19 | 19.91 | |||||||||||||||||||||||
qBRLW7.2 | M488-M489 | 4.71 | -0.19 | 15.17 | |||||||||||||||||||||||
qBRT1 | M66-M67 | 5.70 | -0.05 | 12.94 | |||||||||||||||||||||||
qBRT2 | M170-M171 | 4.45 | -0.04 | 10.26 | |||||||||||||||||||||||
qBRT7.2 | M485-M486 | 5.09 | 0.05 | 11.41 | 4.33 | 0.06 | 13.69 | ||||||||||||||||||||
qBRT7.1 | M488-M489 | 4.59 | 0.07 | 16.39 | |||||||||||||||||||||||
qBRT8 | M544-M545 | 4.98 | -0.05 | 11.25 | |||||||||||||||||||||||
qBRTW1 | M66-M67 | 3.84 | -1.35 | 9.09 | |||||||||||||||||||||||
qBRTW2 | M153-M154 | 3.02 | 1.07 | 7.01 | |||||||||||||||||||||||
qBRTW3 | M204-M205 | 4.70 | -1.41 | 9.58 | 3.16 | -1.54 | 9.90 | ||||||||||||||||||||
qBRTW5.1 | M324-M325 | 6.81 | 1.24 | 14.48 | |||||||||||||||||||||||
qBRTW8.2 | M537-M538 | 5.36 | -1.86 | 13.03 | |||||||||||||||||||||||
qBRTW8.1 | M539-M540 | 4.27 | -1.17 | 8.79 | 4.77 | -1.38 | 15.64 | ||||||||||||||||||||
qBRTW11.1 | M652-M653 | 4.84 | -0.72 | 9.89 |
染色体 Chr. | 区间 Marker interval | 多效性QTL Pleiotropic QTL | 已克隆基因 Cloned gene | 参考文献 Reference |
---|---|---|---|---|
1 | M66-M67 | qBRTW1, qBRT1 | PGL2 | Seonghoe et al. [ |
2 | M123-M124 | qGL2, qBRL2 | GW2 | Song et al. [ |
M136-M137 | qLW2.2, qBRLW2.2 | GS2 | Hu et al. [ | |
M153-M154 | qTGW2, qBRTW2 | |||
3 | M204-M205 | qGL3.1, qLW3.2, qTGW3, qBRTW3, qBRL3.1 | GS3 | Liu et al. [ |
M217-M218 | qGL3.2, qLW3.1, qBRL3.2, qBRLW3.1 | |||
M220-M221 | qGL3.3, qLW3.3, qBRLW3.2 | PGL1 | Dany H et al. [ | |
4 | M245-M246 | qGL4.1, qLW4.1, qBRL4.1 | GL4 | Wu et al. [ |
M255-M256 | qGL4.2, qLW4.2 | |||
5 | M328-M329 | qGW5.1, qLW5, qBRW5.2 | gw5 | Wan et al. [ |
M324-M325 | qBRW5.1, qBRLW5, qBRTW5.1 | |||
7 | M491-M492 | qGW7, qLW7 | GW7 | Wang et al. [ |
M488-M489 | qGT7.1, qBRW7, qBRLW7.2, qBRT7.1 | |||
8 | M537-M538 | qGW8, qGT8.2, qBRL8.1, qBRW8.2, qTGW8.2, qBRTW8.2 | GW8 | Wang et al. [ |
M539-M540 | qBRTW8.1, qGT8.3 | |||
M540-M541 | qTGW8.1, qGT8.1 |
表3 粒重和粒型QTL多效性区域分析
Table 3 Pleiotropic regional analysis of QTLs for grain shape and grain weight.
染色体 Chr. | 区间 Marker interval | 多效性QTL Pleiotropic QTL | 已克隆基因 Cloned gene | 参考文献 Reference |
---|---|---|---|---|
1 | M66-M67 | qBRTW1, qBRT1 | PGL2 | Seonghoe et al. [ |
2 | M123-M124 | qGL2, qBRL2 | GW2 | Song et al. [ |
M136-M137 | qLW2.2, qBRLW2.2 | GS2 | Hu et al. [ | |
M153-M154 | qTGW2, qBRTW2 | |||
3 | M204-M205 | qGL3.1, qLW3.2, qTGW3, qBRTW3, qBRL3.1 | GS3 | Liu et al. [ |
M217-M218 | qGL3.2, qLW3.1, qBRL3.2, qBRLW3.1 | |||
M220-M221 | qGL3.3, qLW3.3, qBRLW3.2 | PGL1 | Dany H et al. [ | |
4 | M245-M246 | qGL4.1, qLW4.1, qBRL4.1 | GL4 | Wu et al. [ |
M255-M256 | qGL4.2, qLW4.2 | |||
5 | M328-M329 | qGW5.1, qLW5, qBRW5.2 | gw5 | Wan et al. [ |
M324-M325 | qBRW5.1, qBRLW5, qBRTW5.1 | |||
7 | M491-M492 | qGW7, qLW7 | GW7 | Wang et al. [ |
M488-M489 | qGT7.1, qBRW7, qBRLW7.2, qBRT7.1 | |||
8 | M537-M538 | qGW8, qGT8.2, qBRL8.1, qBRW8.2, qTGW8.2, qBRTW8.2 | GW8 | Wang et al. [ |
M539-M540 | qBRTW8.1, qGT8.3 | |||
M540-M541 | qTGW8.1, qGT8.1 |
图4 水稻粒型性状QTL对应染色体片段置换系与背景亲本9311的表型差异比较
Fig. 4. Differences of phenotypic values of rice grain shape traits between genetic background parent 9311 and the CSSLs harboring the QTL alleles.
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