\[1\]Garner W W, Allard H A. Effect of the relative length of day and night and other factors of the environment on growth and reproduction in plants. Agric Res, 1920, 18: 553606.\[2\]Garner W W, Allard H A. Further studies in photoperiodism, the response of the plant to relative length of day and night. Agric Res, 1923, 23: 871920.\[3\]种康,雍伟东,谭克辉. 高等植物春化作用研究进展. 植物学通报, 1999, 41(1): 3439.\[4\]Lin H X, Yamamoto T, Sasaki T, et al. Characterization and detection of epistatic interactions of three QTLs, Hd1, Hd2, and Hd3, controlling heading date in rice using nearly isogenic lines. Theor Appl Genet, 1998, 101: 10211028.\[5\]Lin H X, Yamamoto T, Sasaki T, et al. Identification and characterization of a quantitative trait locus, Hd9, controlling heading date in rice. Breed Sci, 2002, 52: 3541.\[6\]Lin H X, Liang Z W, Sasaki T, et al. Fine mapping and characterization of quantitative trait loci Hd4 and Hd5 controlling heading date in rice. Breed Sci, 2003, 53: 5159.\[7\]Lin S Y, Sasaki T, Yano M. Mapping quantitative trait loci controlling seed dormancy and heading date in rice,Oryza sativa L., using backcross inbred lines. Theor Appl Genet, 1997, 96: 9971003.\[8\]Yano M. Genetic and molecular dissection of naturally occurring variation.Curr Opin Plant Biol, 2001, 4: 130135.\[9\]Yano M, Harushima Y, Nagamura Y, et al. Identification of quantitative trait loci controlling heading date in rice using a high density linkage map. Theor Appl Genet, 1997, 95: 10251032.\[10\]Yano M, Katayose Y, Ashikari M, et al. Hd1, a major photoperiod sensitivity quantitative trait locus in rice, is closely related to the Arabidopsis flowering time gene CONSTANS. Plant Cell, 2000, 12: 24732483.\[11\]Yamamoto T,Kuboki Y, Lin S Y, et al. Fine mapping of quantitative trait loci Hd1, Hd2, and Hd3, controlling heading date of rice, as single Mendelian factors. Theor Appl Genet, 1998, 97: 3744.\[12\]Yamamoto T, Lin H X, Sasaki T, et al. Identification of heading date quantitative trait locus Hd6 and characterization of its epistatic interactions with Hd2 in rice using advanced backcross progeny. Genetics, 2000, 154: 885891.\[13\]Kojima S, Takahashi Y, Kobayashi Y, et al. Hd3a, a rice ortholog of the Arabidopsis FT gene, promotes transition to flowering downstream of Hd1 under shortday conditions. Plant Cell Physiol, 2002, 43: 10961105.\[14\]Takahashi Y,Shomura A, Sasaki T, et al. Hd6, a rice quantitative trait locus involved in photoperiod sensitivity, encodes thea subunit of protein kinase CK2. PNAS, 2001, 98: 79227927.\[15\]Luan W, Chen H, Fu Y, et al. The effect of the crosstalk between photoperiod and temperature on the headingdate in rice. PLoS ONE, 2009, 4(6): e5891.\[16\]Komiya R, Ikegami A, Tamaki S, et al. Hd3a and RFT1 are essential for flowering in rice. Development, 2008, 135: 767774.\[17\]Doi K, Izawa T, Fuse T, et al. Ehd1, a Btype response regulator in rice, confers shortday promotion of flowering and controls FTlike gene expression independently of Hd1. Genes Dev, 2004, 18: 926936.\[18\]Hayama R, Yokoi S, Tamaki S, et al. Adaptation of photoperiodic control pathways produces shortday flowering in rice. Nature, 2003, 422: 719722.\[19\]Kim S L, Lee S, Kim H J, et al. OsMADS51 is a shortday flowering promoter that functions upstream of Ehd1, OsMADS14, and Hd3a. Plant Physiol, 2007, 145: 14841494.\[20\]Wu C, You C, Li C, et al. RID1, encoding a Cys2/His2type zinc finger transcription factor, acts as a master switch from vegetative to floral development in rice. PNAS, 2008, 105: 1291512920.\[21\]Matsubara K, Yamanouchi U, Wang X, et al. Ehd2, a rice ortholog of the maize INDETERMINATE1 gene, promotes flowering by upregulating Ehd1. Plant Physiol, 2008, 148: 14251435.\[22\]Park S J, Kim S L, Lee S,et al. Rice indeterminate 1 (OsId1) is necessary for the expression of Ehd1 (early heading date 1) regardless of photoperiod. Plant J, 2008, 56: 10181029.\[23\]Matsubara K, Yamanouchi U, Nonoue Y, et al.Ehd3, encoding a plant homeodomain fingercontaining protein, is a critical promoter of rice flowering. Plant J, 2011, 66: 603612.\[24\]Izawa T, Oikawa T, Tokutomi S, et al. Phytochromes confer the photoperiodic control of flowering in rice (a short day plant). Plant J, 2000, 22: 391399.\[25\]Andre′s F, Galbraith D W, Talo′n M, et al. Analysis of PHOTOPERIOD SENSITIVITY 5 sheds light on the role of phytochromes in photoperiodic flowering in rice. Plant Physiol, 2009, 151: 681690.\[26\]Komiya R, Yokoi R, Shimamoto K. A gene network for longday flowering activates RFT1 encoding a mobile flowering signal in rice. Development, 2009, 136: 34433450.\[27\]Xue W, Xing Y, Weng X, et al. Natural variation in Ghd7 is an important regulator of heading date and yield potential in rice. Nat Genet, 2008, 40: 761767.\[28\]Wei X, Xu J, Guo H, et al. DTH8 suppresses flowering in rice, influencing plant height and yield potential simultaneously. Plant Physiol, 2010, 153:17471758.\[29\]Yan W, Wang P, Chen H, et al. A major QTL, Ghd8, plays pleiotropic roles in regulating grain productivity, plant height, and heading hate in rice. Mol Plant, 2010, 112.\[30\]Hamner K C, Bonner J. Photoperiodism in relation to hormones as factors in floral initiation and development. Bot Gaz, 1938, 100: 388431.\[31\]Ishikawa R, Tamaki S, Yokoi S, et al. Suppression of the floral activator Hd3a is the principal cause of the night break effect in rice. Plant Cell, 2005, 17: 33263336.\[32\]Itoh H, Nonoue Y, Yano M, et al. A pair of floral regulators sets critical day length for Hd3a florigen expression in rice. Nat Genet, 2010, 42(7): 635639.\[33\]Izawa T,OikawaT, Sugiyama N, et al. Phytochrome mediates the external light signal to repress FT orthologs in photoperiodic flowering of rice. Genes Dev, 2002, 16: 2006 2020.\[34\]Takano M, Inagaki N,Xie X, et al. Distinct and cooperative functions of phytochromes A, B, and C in the control of deetiolation and flowering in rice. Plant Cell, 2005, 17: 33113325.\[35\]Takano M, Inagaki N, Xie X, et al. Phytochromes are the sole photoreceptors for perceiving red/farred light in rice. PNAS, 2009, 106(34): 1470514710.\[36\]Ishikawa R, Aoki M, Kurotani K, et al. Phytochrome B regulates heading date 1(Hd1)mediated expression of rice florigen Hd3a and critical day length in rice. Mol Genet Genom, 2011, 285: 461470.\[37\]Zhang Y, Gong S, Li Q, et al. Functional and signaling mechanism analysis of rice CRYPTOCHROME 1. Plant J, 2006, 46: 971983.\[38\]Hirose F, Shinomura T, Tanabata T et al. Involvement of rice cryptochromes in deetiolation responses and flowering. Plant Cell Physiol, 2006, 47 (7): 915925.\[39\]Michaels S D,Amasino R M. FLOWERING LOCUS C encodes a novel MADS domains protein that acts as a repressor of flowering. Plant Cell, 1999, 11:949956.\[40\]Searle I, He Y H,Turck F, et al. The transcription factor FLC confers a flowering response to verbalization by repressing meristem competence and systemic signaling in Arabidopsis. Genes & Dev, 2006, 20: 898912.\[41\]Blázquez M A, Ahn J H, Weigel D. A thermosensory pathway controlling flowering time in Arabidopsis thaliana. Nature Genet, 2003, 33: 168171.\[42\]Yan L,Loukoianov A, Tranquilli G, et al. Positional cloning of the wheat vernalization gene VRN1. PNAS, 2003, 100: 62636268.\[43\]Yan L, Loukoianov A, Blechl A, et al. The wheat VRN2 gene is a flowering repressor downregulated by vernalization. Science, 2004, 303: 16401644.\[44\]Yan L, Fu D, Li C,et al. The wheat and barley vernalization gene VRN3 is an orthologue of FT. PNAS, 2006, 103: 1958119586.\[45\]Sheldon C C, Burn E J, Perez P P, et al. The FLF MADS box gene: A repressor of flowering in Arabidopsis regulated by vernalization and methylation. Plant Cell, 1999, 11:445458.\[46\]Sheldon C C, Rouse D T, Finnegan E J, et al. The molecular basis of vernalization: The central role of FLOWERING LOCUS C (FLC). PNAS, 2000, 97: 37533758.\[47\]Lee J H,Yoo S J, Park S H, et al. Role of SVP in the control of flowering time by ambient temperature in Arabidopsis. Genes Dev, 2007, 21: 397402.\[48\]Trevaskis B, Tadege M, Hemming M N, et al. Short vegetative phaselike MADS box genes inhibit floral meristem identity in barley. Plant Physiol, 2007, 143: 225235.\[49\]Lee S, Choi S C, An G. Rice SVPgroup MADSbox proteins,OsMADS22 and OsMADS55,are negative regulators of brassinosteroid responses. Plant J, 2008, 54: 93105. |